In ecology, shade tolerance is a plant's ability to tolerate low light levels. The term is also used in horticulture and landscaping, although in this context its use is sometimes imprecise, especially in labeling of plants for sale in commercial nurseries. Shade tolerance is a complex, multi-faceted property of plants. Different plant species exhibit different adaptations to shade, and a particular plant can exhibit varying degrees of shade tolerance, or even of requirement for light, depending on its history or stage of development. Basic concepts Except for some parasitic plants, all land plants need sunlight to survive. However, in general, more sunlight does not always make it easier for plants to survive. In direct sunlight, plants face desiccation and exposure to UV rays, and must expend energy producing pigments to block UV light, and waxy coatings to prevent water loss. Plants adapted to shade have the ability to use far-red light (about 730 nm) more effectively than plants adapted to full sunlight. Most red light gets absorbed by the shade-intolerant canopy plants, but more of the far-red light penetrates the canopy, reaching the understorey. The shade-tolerant plants found here are capable of photosynthesis using light at such wavelengths. The situation with respect to nutrients is often different in shade and sun. Most shade is due to the presence of a canopy of other plants, and this is usually associated with a completely different environment—richer in soil nutrients—than sunny areas. Shade-tolerant plants are thus adapted to be efficient energy-users. In simple terms, shade-tolerant plants grow broader, thinner leaves to catch more sunlight relative to the cost of producing the leaf. Shade-tolerant plants are also usually adapted to make more use of soil nutrients than shade-intolerant plants. A distinction may be made between "shade-tolerant" plants and "shade-loving" or sciophilous plants. Sciophilous plants are dependent on a degree of shading that would eventually kill most other plants, or significantly stunt their growth. Plants adaptation to the changing light Plants applied multilevel adaptations to the changing light environment from the systemic level to the molecular level. Leaf movement Various types of leaf movement for adaptation to changing light environment have been identified: developmental, passive and active. Active movements are reversible. Some plants use blue-light absorbing pigments as a sensor and pulvinar motor tissue to drive leaf movement. These adaptions are usually slow but relatively efficient. They are advantageous to some shade plant that have