with 1.4%, and Ecuador with 1%. Natural The rainforest likely formed during the Eocene era (from 56 million years to 33.9 million years ago). It appeared following a global reduction of tropical temperatures when the Atlantic Ocean had widened sufficiently to provide a warm, moist climate to the Amazon basin. The rainforest has been in existence for at least 55 million years, and most of the region remained free of savanna-type biomes at least until the current ice age when the climate was drier and savanna more widespread. Following the Cretaceous–Paleogene extinction event, the extinction of the dinosaurs and the wetter climate may have allowed the tropical rainforest to spread out across the continent. From 66 to 34 Mya, the rainforest extended as far south as 45°. Climate fluctuations during the last 34 million years have allowed savanna regions to expand into the tropics. During the Oligocene, for example, the rainforest spanned a relatively narrow band. It expanded again during the Middle Miocene, then retracted to a mostly inland formation at the last glacial maximum. However, the rainforest still managed to thrive during these glacial periods, allowing for the survival and evolution of a broad diversity of species. During the mid-Eocene, it is believed that the drainage basin of the Amazon was split along the middle of the continent by the Púrus Arch. Water on the eastern side flowed toward the Atlantic, while to the west water flowed toward the Pacific across the Amazonas Basin. As the Andes Mountains rose, however, a large basin was created that enclosed a lake; now known as the Solimões Basin. Within the last 5–10 million years, this accumulating water broke through the Púrus Arch, joining the easterly flow toward the Atlantic. There is evidence that there have been significant changes in the Amazon rainforest vegetation over the last 21,000 years through the last glacial maximum (LGM) and subsequent deglaciation. Analyses of sediment deposits from Amazon basin paleolakes and the Amazon Fan indicate that rainfall in the basin during the LGM was lower than for the present, and this was almost certainly associated with reduced moist tropical vegetation cover in the basin. In present day, the Amazon receives approximately 9 feet of rainfall annually. There is a debate, however, over how extensive this reduction was. Some scientists argue that the rainforest was reduced to small, isolated refugia separated by open forest and grassland; other scientists