long time, it was thought that the Amazon rainforest was never more than sparsely populated, as it was impossible to sustain a large population through agriculture given the poor soil. Archeologist Betty Meggers was a prominent proponent of this idea, as described in her book Amazonia: Man and Culture in a Counterfeit Paradise. She claimed that a population density of is the maximum that can be sustained in the rainforest through hunting, with agriculture needed to host a larger population. However, recent anthropological findings have suggested that the region was actually densely populated. Some 5 million people may have lived in the Amazon region in AD 1500, divided between dense coastal settlements, such as that at Marajó, and inland dwellers. By 1900, the population had fallen to 1 million and by the early 1980s it was less than 200,000. The first European to travel the length of the Amazon River was Francisco de Orellana in 1542. The BBC's Unnatural Histories presents evidence that Orellana, rather than exaggerating his claims as previously thought, was correct in his observations that a complex civilization was flourishing along the Amazon in the 1540s. It is believed that civilization was later devastated by the spread of diseases from Europe, such as smallpox. This civilization was investigated by the British explorer Percy Fawcett in the early twentieth century. The results of his expeditions were inconclusive and he disappeared mysteriously on his last trip. His name for this lost civilization was the City of Z. Since the 1970s, numerous geoglyphs have been discovered on deforested land dating between AD 1–1250, furthering claims about Pre-Columbian civilizations. Ondemar Dias is accredited with first discovering the geoglyphs in 1977, and Alceu Ranzi is credited with furthering their discovery after flying over Acre. The BBC's Unnatural Histories presented evidence that the Amazon rainforest, rather than being a pristine wilderness, has been shaped by man for at least 11,000 years through practices such as forest gardening and terra preta. Terra preta is found over large areas in the Amazon forest; and is now widely accepted as a product of indigenous soil management. The development of this fertile soil allowed agriculture and silviculture in the previously hostile environment; meaning that large portions of the Amazon rainforest are probably the result of centuries of human management, rather than naturally occurring as has previously been supposed. In the region of the Xingu tribe, remains of