Otters are carnivorous mammals in the subfamily Lutrinae. The 13 extant otter species are all semiaquatic, aquatic, or marine, with diets based on fish and invertebrates. Lutrinae is a branch of the Mustelidae family, which also includes weasels, badgers, mink, and wolverines, among other animals. Etymology The word otter derives from the Old English word or . This, and cognate words in other Indo-European languages, ultimately stem from the Proto-Indo-European language root , which also gave rise to the English word "water". Terminology An otter's den is called a holt or couch. Male otters are called dogs or boars, females are called bitches or sows, and their offspring are called pups or cubs. The collective nouns for otters are bevy, family, lodge, romp (being descriptive of their often playful nature) or, when in water, raft. The feces of otters are typically identified by their distinctive aroma, the smell of which has been described as ranging from freshly mown hay to putrefied fish; these are known as spraints. Life cycle The gestation period in otters is about 60 to 86 days. The newborn pup is cared for by the bitch, dog, and older offspring. Female otters reach sexual maturity at approximately two years of age and males at approximately three years. The holt is built under tree roots or a rocky cairn, more common in Scotland. It is lined with moss and grass. After one month, the pup can leave the holt and after two months, it is able to swim. The pup lives with its family for approximately one year. Otters live up to 16 years; they are by nature playful, and frolic in the water with their pups. Its usual source of food is fish, and further downriver, eels, but it may sample frogs and birds. Description Otters have long, slim bodies and relatively short limbs. Their most striking anatomical features are the powerful webbed feet used to swim, and their seal-like abilities for holding breath underwater. Most have sharp claws on their feet and all except the sea otter have long, muscular tails. The 13 species range in adult size from in length and in weight. The Asian small-clawed otter is the smallest otter species and the giant otter and sea otter are the largest. They have very soft, insulated underfur, which is protected by an outer layer of long guard hairs. This traps a layer of air