cases, a coalition will consist of brothers born in the same litter who stayed together after weaning, but biologically unrelated males are often allowed into the group; in the Serengeti 30% members in coalitions are unrelated males. If a cub is the only male in a litter he will typically join an existing group, or form a small group of solitary males with two or three other lone males who may or may not be territorial. In the Kalahari Desert around 40% of the males live in solitude. Males in a coalition are affectionate toward each other, grooming mutually and calling out if any member is lost; unrelated males may face some aversion in their initial days in the group. All males in the coalition typically have equal access to kills when the group hunts together, and possibly also to females who may enter their territory. A coalition generally has a greater chance of encountering and acquiring females for mating, however, its large membership demands greater resources than do solitary males. A 1987 study showed that solitary and grouped males have a nearly equal chance of coming across females, but the males in coalitions are notably healthier and have better chances of survival than their solitary counterparts. Home ranges and territories Unlike many other felids, among cheetahs, females tend to occupy larger areas compared to males. Females typically disperse over large areas in pursuit of prey, but they are less nomadic and roam in a smaller area if prey availability in the area is high. As such, the size of their home range depends on the distribution of prey in a region. In central Namibia, where most prey species are sparsely distributed, home ranges average , whereas in the woodlands of the Phinda Game Reserve (South Africa), which have plentiful prey, home ranges are in size. Cheetahs can travel long stretches overland in search of food; a study in the Kalahari Desert recorded an average displacement of nearly every day and walking speeds ranged between . Males are generally less nomadic than females; often males in coalitions (and sometimes solitary males staying far from coalitions) establish territories. Whether males settle in territories or disperse over large areas forming home ranges depends primarily on the movements of females. Territoriality is preferred only if females tend to be more sedentary, which is more feasible in areas with plenty of prey. Some males,