settlers from the east, and were obliged to accept the dominance of Carthage for centuries. Nonetheless, therein they persisted largely unassimilated, as a separate, submerged entity, as a culture of mostly passive urban and rural poor within the civil structures created by Punic rule. In addition, and most importantly, the Berber peoples also formed quasi-independent satellite societies along the steppes of the frontier and beyond, where a minority continued as free 'tribal republics'. While benefiting from Punic material culture and political-military institutions, these peripheral Berbers (also called Libyans)—while maintaining their own identity, culture, and traditions—continued to develop their own agricultural skills and village societies, while living with the newcomers from the east in an asymmetric symbiosis. As the centuries passed, there naturally grew a Punic society of Phoenician-descent but born in Africa, called . This term later came to be applied also to Berbers acculturated to urban Phoenician culture. Yet the whole notion of a Berber apprenticeship to the Punic civilization has been called an exaggeration sustained by a point of view fundamentally foreign to the Berbers. There evolved a population of mixed ancestry, Berber and Punic. There would develop recognized niches in which Berbers had proven their utility. For example, the Punic state began to field Berber–Numidian cavalry under their commanders on a regular basis. The Berbers eventually were required to provide soldiers (at first "unlikely" paid "except in booty"), which by the fourth century BC became "the largest single element in the Carthaginian army". Yet in times of stress at Carthage, when a foreign force might be pushing against the city-state, some Berbers would see it as an opportunity to advance their interests, given their otherwise low status in Punic society. Thus, when the Greeks under Agathocles (361–289 BC) of Sicily landed at Cape Bon and threatened Carthage (in 310 BC), there were Berbers, under Ailymas, who went over to the invading Greeks. During the long Second Punic War (218–201 BC) with Rome (see below), the Berber King Masinissa ( BC) joined with the invading Roman general Scipio, resulting in the war-ending defeat of Carthage at Zama, despite the presence of their renowned general Hannibal; on the other hand, the Berber King Syphax (d. 202 BC) had supported Carthage. The Romans, too, read these cues, so that they cultivated their Berber alliances and, subsequently, favored the Berbers who advanced their interests following the Roman victory. Carthage was faulted