The even-toed ungulates (Artiodactyla , ) are ungulates—hoofed animals—which bear weight equally on two (an even number) of their five toes: the third and fourth. The other three toes are either present, absent, vestigial, or pointing posteriorly. By contrast, odd-toed ungulates bear weight on an odd number of the five toes. Another difference between the two is that many other even-toed ungulates (with the exception of Suina) digest plant cellulose in one or more stomach chambers rather than in their intestine as the odd-toed ungulates do. The advent of molecular biology, along with new fossil discoveries, found that Cetaceans (whales, dolphins, and porpoises) fall within this taxonomic branch, being most closely related to hippopotamuses. Some modern taxonomists thus apply the name Cetartiodactyla to this group, while others opt to include cetaceans within the existing name of Artiodactyla. The roughly 270 land-based even-toed ungulate species include pigs, peccaries, hippopotamuses, antelopes, deer, giraffes, camels, llamas, alpacas, sheep, goats, and cattle. Many are herbivores, but suids are omnivorous, while cetaceans are almost exclusively carnivorous. Many of these are of great dietary, economic, and cultural importance to humans. Evolutionary history The oldest fossils of even-toed ungulates date back to the early Eocene (about 53 million years ago). Since these findings almost simultaneously appeared in Europe, Asia, and North America, it is very difficult to accurately determine the origin of artiodactyls. The fossils are classified as belonging to the family Dichobunidae; their best-known and best-preserved member is Diacodexis. These were small animals, some as small as a hare, with a slim build, lanky legs, and a long tail. Their hind legs were much longer than their front legs. The early to middle Eocene saw the emergence of the ancestors of most of today's mammals. Two formerly widespread, but now extinct, families of even-toed ungulates were Entelodontidae and Anthracotheriidae. Entelodonts existed from the middle Eocene to the early Miocene in Eurasia and North America. They had a stocky body with short legs and a massive head, which was characterized by two humps on the lower jaw bone. Anthracotheres had a large, porcine (pig-like) build, with short legs and an elongated muzzle. This group appeared in the middle Eocene up until the Pliocene, and spread throughout Eurasia, Africa, and North America. Anthracotheres are thought to be the ancestors of hippos, and, likewise, probably led a similar aquatic lifestyle. Hippopotamuses appeared in the late Miocene and occupied Africa