better born ...." He lists other ethnic populations among the settlers: Abantes from Euboea, Minyans from Orchomenus, Cadmeians, Dryopians, Phocians, Molossians, Arcadian Pelasgians, Dorians of Epidaurus, and others. Even "the best born of the Ionians" had married girls from Caria. He defines Ionians as all peoples who were descended from Athenians and celebrated the Apaturia festival, which aligns with the expansive Athenian definition of Ionian identity. Satrapy (387–335 BC) The Spartans dissolved the Athenian Empire at the end of the Peloponnesian War in 404 BC. The Spartans installed harmosts (governors) in the cities, but had to withdraw them because they had promised Ionia and the other Greek communities in Asia to the Persians. In 401, the Ionian cities and Sparta supported Cyrus the Younger, the Persian overlord of Asia Minor, in his attempt to seize the throne from his brother, King Artaxerxes II but he failed. Artaxerxes tasked Tissaphernes, the satrap of Lydia and Caria, with retaking the Ionian cities, and the Spartans opposed him. In 396 BC, Agesilaus led a large expedition to Asia Minor to defend the cities and attack the Persians, which landed in Ephesus. From there he invaded Phrygia and Lydia, sacking Sardis in 395 BC. But the outbreak of the Corinthian War forced him to withdraw in 395 BC. The region was under Persian control by about 390 BC, when the Persian satrap arbitrated a boundary dispute between Miletus and Myus. Sparta, Athens, and the other mainland Greek states formally acknowledged Persian possession of Ionia and the other Greek cities in Asia Minor in the Peace of Antalcidas in 387 BC. In this period, Ionia was a separate satrapy, rather than part of Lydia - the only time in the region's history that formed an administrative unit. Ionian cities appear to have retained a considerable amount of autonomy until the conquest of Asia Minor by Alexander the Great in 335 BC. Hellenistic period Ephesus was conquered by Philip II of Macedon in 336 BC in preparation for the invasion of Persia, which took place under his son Alexander the Great. After the battle of the Granicus in 334 BC most of the Ionian cities submitted to Alexander, except for Miletus, which was taken only after a long siege. Alexander presented his invasion as a liberation of the Greeks of Asia and therefore treated the Ionians generously, granting them freedom, autonomy, and tax-free status. In the