show evidence that they attached to the ankle, according to Wang et al.. They are relatively elongated with a wingspan of ninety centimetres. The legs are short but robust. The toes bear well-developed curved claws, but these are not as long as the hand claws. The fifth toe is elongated, according to the authors supporting a membrane between the legs, the uropatagium. Phylogeny Jeholopterus was by the authors assigned to the Anurognathidae. An analysis by Lü Junchang in 2006 resolved its position as being the sister taxon of Batrachognathus. Kellner and colleagues in 2009 proposed Batrachognathinae for the clade comprising these two genera and Dendrorhynchoides. In 2021, a phylogenetic analysis conducted by Xuefang Wei and colleagues recovered Jeholopterus within the subfamily Anurognathinae, which is unlike the former study by Kellner and colleagues where it was recovered as closely related to Batrachognathus. Jeholopterus was found to have been the sister taxon to an unnamed clade formed by both Anurognathus and Vesperopterylus in Wei and colleagues' study. Below is a cladogram representing their phylogenetic analysis: Lifestyle Anurognathids are normally considered insectivores. Wang et al hypothesized that Jeholopterus, being the largest species known of the group, might also have been a piscivore, a fish-eater. See also List of pterosaur genera Timeline of pterosaur research References Monofenestratans Middle Jurassic pterosaurs of Asia Late Jurassic pterosaurs of Asia Taxa named by Zhou Zhonghe Fossil taxa described in 2002 Taxa named by Xu Xing Paleontology in Inner Mongolia