concentrate in the canopy and spread continuously if the shrubs are close enough together. Shrublands are typically dry and are prone to accumulations of highly volatile fuels, especially on hillsides. Fires will follow the path of least moisture and the greatest amount of dead fuel material. Surface and below-ground soil temperatures during a burn are generally higher than those of forest fires because the centers of combustion lie closer to the ground, although this can vary greatly. Common plants in shrubland or chaparral include manzanita, chamise and coyote brush. California shrublands California shrubland, commonly known as chaparral, is a widespread plant community of low growing species, typically on arid sloping areas of the California Coast Ranges or western foothills of the Sierra Nevada. There are a number of common shrubs and tree shrub forms in this association, including salal, toyon, coffeeberry and Western poison oak. Regeneration following a fire is usually a major factor in the association of these species. South African Fynbos shrublands Fynbos shrublands occur in a small belt across South Africa. The plant species in this ecosystem are highly diverse, yet the majority of these species are obligate seeders, that is, a fire will cause germination of the seeds and the plants will begin a new life-cycle because of it. These plants may have coevolved into obligate seeders as a response to fire and nutrient-poor soils. Because fire is common in this ecosystem and the soil has limited nutrients, it is most efficient for plants to produce many seeds and then die in the next fire. Investing a lot of energy in roots to survive the next fire when those roots will be able to extract little extra benefit from the nutrient-poor soil would be less efficient. It is possible that the rapid generation time that these obligate seeders display has led to more rapid evolution and speciation in this ecosystem, resulting in its highly diverse plant community. Grasslands Grasslands burn more readily than forest and shrub ecosystems, with the fire moving through the stems and leaves of herbaceous plants and only lightly heating the underlying soil, even in cases of high intensity. In most grassland ecosystems, fire is the primary mode of decomposition, making it crucial in the recycling of nutrients. In some grassland systems, fire only became the primary mode of decomposition after the disappearance of large migratory herds of browsing or grazing megafauna driven