Brocchinia is a genus of the botanical family Bromeliaceae, and is the sole genus of the subfamily Brocchinioideae, containing 20 species. The genus is named for Giovanni Battista Brocchi, Italian naturalist (1772–1826). Brocchinia species are native primarily to the ancient Guayana Shield in southern Venezuela and Guyana, with some species extending into Colombia and northern Brazil. Its species are generally restricted to areas of sand and sandstone of the Roraima Formation; a few occur on granite. Based on chloroplast DNA sequence variation, Brocchinia appears to be sister to all other bromeliads. Calibration of the molecular family tree of bromeliads against the known ages of various fossil monocots suggests that Brocchinia lineage diverged from other bromeliads nearly 20 million years ago, and that some of the living species of Brocchinia began diverging from each other soon thereafter. The next lineage to diverge from other bromeliads included Lindmania and Connellia, which are also endemic to the Guayana Shield; at least one other subfamily Navioideae is also almost entirely restricted to that region. Consequently, it is quite likely that the family as a whole arose in the Guayana Shield. Previously, Brocchinia was thought to be a member of the bromeliad subfamily Pitcairnioideae, based on its possession of winged seeds like those seen in other members of that group. This view has now been overturned after the recognition that Pitcairnioideae as originally circumscribed more than a century ago represents several morphologically similar but independently derived groups that share only ancestral (plesiomorphic) characters, not true synapomorphies (shared derived characters). Brocchinia has undergone a spectacular adaptive radiation in mechanisms of nutrient capture, apparently in response to the unusually infertile, heavily leached substrates of the Guayana Shield. At least two species, Brocchinia reducta and B. hechtioides, appear to be carnivorous. Like most species of Brocchinia – and indeed, of many other bromeliad genera – these two species impound rain water in a tank formed by tightly overlapping leaves arranged in a basal rosette. In these species, however, the leaves are nearly vertical, their inner surfaces are covered with fine wax that readily exfoliates, the tank fluid is highly acid (pH 3.0) and emits a sweet, nectar-like odor, and the tank itself is full of the dead remains of ants (B. reducta) or bees and wasps (B. hechtioides). Recently, it has been shown that at least B. reducta secretes a phosphatase into its tanks. Another species, Brocchinia acuminata,