the brain that are sensitive to specific olfactory receptors. He holds the titles of University Professor at Columbia University, Professor of Biochemistry and Molecular Biophysics and of Pathology at Columbia University's College of Physicians and Surgeons, and Investigator of the Howard Hughes Medical Institute. In addition to contributions to neurobiology, Axel has also made seminal discoveries in immunology, and his lab was one of the first to identify the link between HIV infection and immunoreceptor CD4. In addition to making contributions as a scientist, Axel has also mentored many leading scientists in the field of neurobiology. Seven of his trainees have become members of the National Academy of Sciences, and currently six of his trainees are affiliated with the Howard Hughes Medical Institute's investigator and early scientist award programs. Awards and honors In addition to the Nobel Prize, Axel has won numerous awards and honors. He was elected a Fellow of the American Academy of Arts and Sciences and a member of the National Academy of Sciences in 1983. In 2005, Axel received the Golden Plate Award of the American Academy of Achievement. Axel was awarded the Double Helix Medal in 2007. CSHL Double Helix Medal Honoree and was elected a Foreign Member of the Royal Society (ForMemRS) in 2014. His nomination reads: Personal life Axel is married to fellow scientist and olfaction pioneer Cornelia Bargmann. Previously, he had been married to Ann Axel, who is a social worker at Columbia University Medical Center. Owing to his tall stature, Axel played basketball during high school. References External links - pressrelease 1946 births Living people Nobel laureates in Physiology or Medicine American Nobel laureates Richard-Lounsbery Award laureates Members of the United States National Academy of Sciences American neuroscientists Jewish biologists Jewish American scientists Stuyvesant High School alumni Columbia College (New York) alumni Columbia University alumni Johns Hopkins School of Medicine alumni Columbia Medical School faculty Howard Hughes Medical Investigators Fellows of the American Academy of Arts and Sciences Foreign Members of the Royal Society Members of the American Philosophical Society Scientists from New York (state)