rosettes of teeth, as well as a more developed "keeled" crest compared to Ornithocheirus. Another feature that made Ornithocheirus unique and unlike its relatives, was that its teeth of were mostly vertical, rather than set at an outward-pointing angle. It was believed in the past that Ornithocheirus was one of the largest pterosaurs to have existed, with a wingspan possibly measuring 40 feet (12.2 m) wide. However, this is a highly exaggerated number, as the animal's wingspan likely measured 15 to 20 feet (4.5 to 6.1 m) wide, which would make it a medium-sized pterosaur. A related species called Tropeognathus had a wingspan measuring about wide, making it the largest toothed pterosaur known. In 2022, Gregory S. Paul estimated that Ornithocheirus had a wingspan of and a body mass of . Classification A topology made by Andres and Myers in 2013 placed Ornithocheirus within the family Ornithocheiridae in a more derived position than Tropeognathus, but in a more basal position than Coloborhynchus, and the family itself is placed within the more inclusive clade Ornithocheirae. In 2019, Pêgas et al. found Ornithocheirus to be in a basal member of the clade Ornithocheirae, reclassifying all other snout-crested pterosaurs in the family Anhangueridae. Their cladogram is shown on the right. Topology 1: Andres & Myers (2013). Topology 2: Pêgas et al. (2019). See also List of pterosaur genera Timeline of pterosaur research Pterosaur size Notes Haines, T., and Chambers, P. (2006). The Complete Guide to Prehistoric Life''. Canada: Firefly Books Ltd. References Pteranodontoids Late Cretaceous pterosaurs of Europe Late Cretaceous reptiles of Europe Taxa named by Harry Seeley Fossil taxa described in 1869