The Malagasy () are an Austronesian-speaking African ethnic group indigenous to the island country of Madagascar. Traditionally, the population have been divided by subgroups (tribes or ethnicities). Examples include "Highlander" (ethnically Austronesian/Malay race-Indonesian with less Bantu ancestry) groups such as the Merina and Betsileo of the central highlands around Antananarivo, Alaotra (Ambatondrazaka) and Fianarantsoa, and the "coastal dwellers" (ethnically Bantu with less Austronesian ancestry) with tribes like the Sakalava, Bara, Vezo, Betsimisaraka, Mahafaly, etc. The Merina are also further divided into two subgroups. The “Merina A” are the Hova and Andriana, and have an average of 30–40% Bantu ancestry. The second subgroup is the “Merina B”, the Andevo, who have an average of 40–50% Bantu ancestry. They make up less than 1/3 of Merina society. The Malagasy population was 2,242,000 in the first census in 1900. Their population experienced a massive growth in the next hundred years, especially under French Madagascar. Genetics and origins of the populations The genetics element about the dual origin of the Malagasy can be traced to the mid-20th century with results regarding blood group distribution. An expansive island-wide survey of the genetic diversity has been performed from 2008 to 2018. This project was called "MAGE" (for Madagascar, Anthropology Genetics Ethno-linguistic). Around 3000 inhabitants of Madagascar have participated in this study and provided their saliva for a genetic study. Three hundred villages across Madagascar have been sampled in terms of genetic, linguistic and cultural diversity. This research was led and performed by Malagasy and European researchers and academics. This study demonstrated that all Malagasy people have mixed African and Asian ancestry. But the proportion of ancestral genes differs. Coastal Malagasy populations, including the Temoro, Vezo, and Mikea, etc. have approximately 70% African ancestry and 30% Asian ancestry, while highlander tribes tend to have lower African ancestry at around 45%. In a recent island-wide survey the male-only Y chromosomes of African origin are more common than those of East Asian origin, but it varies depending on the study (70.7 vs. 20.7 or 51% vs 34%). However the mtDNA lineages, passed down from mother to child, are the opposite (42.4% African origin vs. 50.1% East Asian origin). Due to the proximity to Africa, the connection with Asian populations exhibited the most curiosity. Around 1996, a study was launched in an attempt to identify the presence of the Polynesian motif in the Malagasy population (mtDNA haplotype B4a1a1a). A more