Gellius and Pliny the Elder both took Boreas as a northeast wind, equivalent to the Roman Aquilo, similar to Nor'easter winter storms. Boreas is depicted as being very strong, with a violent temper to match. He was frequently shown as a winged old man or sometimes as a young man with shaggy hair and beard, holding a conch shell and wearing a billowing cloak. Pausanias wrote that Boreas had snakes instead of feet, though in art he was usually depicted with winged human feet. Boreas was closely associated with horses, storms and winter. He was said to have fathered twelve colts after taking the form of a stallion, to the mares of Erichthonius, king of Dardania. These were said to be able to run across a field of grain without trampling the plants. Pliny the Elder (Natural History iv.35 and viii.67) thought that mares might stand with their hindquarters to the North Wind and bear foals without a stallion. The Greeks believed that his home was in Thrace, and Herodotus and Pliny both describe a northern land known as Hyperborea "Beyond the North Wind" where people lived in complete happiness and had extraordinarily long lifespans. He is said to have fathered three giant Hyperborean priests of Apollo by Chione. When the goddess Leto, pregnant with Artemis and Apollo, was due, Boreas was ordered by Zeus to bring her to Poseidon, who in turn led her to the island of Ogygia where she could give birth to the twins, as Zeus' wife Hera had ordered all places and land to shun Leto. Boreas was also said to have kidnapped Orithyia, an Athenian princess, from the Ilisos. Boreas had taken a fancy to Orithyia and had initially pleaded for her favours, hoping to persuade her. When this failed, he reverted to his usual temper and abducted her as she danced on the banks of the Ilisos. Boreas wrapped Orithyia up in a cloud, raped her, and with her, Boreas fathered two sons—the Boreads, Zethes and Calais, who were part of the crew of the Argo as Argonauts.—and two daughters—Chione, goddess of snow, and Cleopatra. From then on, the Athenians saw Boreas as a relative by marriage. When Athens was threatened by Xerxes, the people prayed to Boreas, who was said to have then caused winds to sink 400 Persian ships. A cult was established in Athens in 480 B. C. E. in