Park, east of Santa Marta. Cebus a. cesarae can be located in the Serranía del Perijá east of Valledupar, Cesar also in Colombia. Behavior and ecology White-fronted capuchins have been studied in Colombia by Defler, in two different sites in Peru by Soini and Terborgh, in Trinidad by Phillips and in Ecuador by Matthews. In eastern Vichada, Colombia, white-fronted capuchins are found in large groups of around 35 individuals, while to the south in closed forest (perhaps as a result of competition with the tufted capuchin) they have an average group size of 8–15 individuals. A group in Vichada used a home range of about , while Terborgh found a home range of more than and Matthews calculated . Near the type locality in gallery forest and islands of forest in Vichada, they have an ecological density of around 30 individuals/km2. In forests with closed-canopy in Colombia and in southern Vichada, many areas have very low densities. Around the lower Apaporis River, for example, densities are less than one individual/km2 and the size of the groups is around 15 individuals. Low densities in many parts of the Colombian Amazon make it difficult to detect the presence of the species in many parts. Terborgh found an average of for the day range of a group, and calculated the following time budget of the study group in Manú National Park, Peru: 18% rest, 21% travel, 22% feeding on plant material and 39% feeding on insects; total feeding 61%. Matthews however, registered 54% foraging, 25% moving and 21% feeding and socializing. They are primarily quadrupedal, although they utilize a great variety of gallops, jumps, falls and climbing. During certain times of the year they are extremely terrestrial, especially when there is a scarcity of available fruits and the troop must search for arthropods in the dry leaves of the forest floor. In some parts of the Llanos Orientales they are found walking over the grassy savanna between forests, leaving well-beaten trails. In Vichada it uses preferential trees for sleeping at heights of . The palm Attalea regia is often used for sleeping in this zone. Diet All species of capuchin tend to have a rather similar diet in broad terms; they are omnivores, eating fruits and small invertebrates, small vertebrates and birds' eggs, which they forage at all levels of the forest, frequently descending to the forest floor. In northern Colombia during the