and colleagues, 2015: Cheke and the Dutch ornithologist Justin J. F. J. Jansen stated in 2016 that the Edinburgh specimen has no clear provenance information, and that it may have been collected from Mauritius (only one of Dufresne's other bird specimens was from Réunion, while several were from Mauritius). They noted that, unlike modern Mauritian specimens, the pink neck ring of the Edinburgh specimen continued uninterrupted around the back of the neck, similar to what Buffon and Levaillant described, but that from where the specimen Levaillant described was, was unclear. They stated that the genetic differences between the specimens were not necessarily subspecific, but because the Mauritian specimens were much more recent than the Edinburgh specimen, the similarity of the former specimens could have been due to a genetic bottleneck, resulting from a severe decline of the Mauritian population in the 19th century. They concluded that the default assumption should be that it came from Réunion. They also called attention to a usually overlooked, unlabelled sketch from around 1770 by French artist Paul Philippe Sanguin de Jossigny of a ring-necked parakeet with a collar encircling the neck, which they thought could have been from either island. In 2017, Hume agreed that the Edinburgh specimen could have come from Mauritius. He stated that the genetic differences could be due to variation within the population there, and pointed out that some other bird species migrate between Mauritius and Réunion. Also in 2017, Australian ornithologist Joseph M. Forshaw agreed that the Mauritius and Réunion populations were subspecifically distinct and that the Edinburgh specimen was from Réunion, and should be designated the neotype of P. eques. The following year, Jones and colleagues, including authors of the DNA studies, Hume, and Forshaw, supported the identification of the Edinburgh specimen as a Réunion parakeet and the subspecific differentiation between the populations. They found that the specimen differed from all examined Mauritius specimens in having a complete pink collar, instead of having a gap at the back of the neck, a feature emphasised by Brisson, Buffon, and Levaillant in their descriptions of the Réunion parakeet, but not obvious in the photographs seen by Jones in the 1980s. Since populations on islands usually have lower genetic diversity than those on continents, they stated that the low level of differentiation between the Mauritius and Réunion specimens would be expected. They also concluded that Jossigny's drawing showed a Réunion parakeet.