European population of the time, now known as Western Hunter-Gatherers, another example being Loschbour man discovered in Luxembourg. This population forms about 10%, on average, of the ancestry of Britons without a recent family history of immigration. Brown eyes, lactose tolerance, and light skin are common in the modern population of the area. These genes came from later immigration, most of it ultimately from two major waves, the first of Neolithic farmers from the Near East, another of Bronze Age pastoralists, most likely speakers of Indo-European languages, from the Pontic steppe. Ancestry About 85% of his ancestry can be modelled as coming from the ~14,000–7,000-year-old Villabruna genetic cluster, and only c. 15% from the Goyet Q2 cave cluster whose genes are found in association with the Late Upper Palaeolithic Magdalenian culture. He is not closely related to the earlier Magdalenian individuals found in the same cave, whose ancestry is entirely from the Goyet cluster. The genomes of all British Mesolithic individuals sequenced to date other than Cheddar Man can be modelled as only Villabruna-related (WHG) ancestry, without additional Goyet-related admixture. Uniparental haplogroups Cheddar Man's Y-DNA belonged to an ancient sister branch of modern I2-L38 (I2a2). The I2a2 subclade is still extant in males of the modern British Isles and across other parts of Europe. The mitochondrial DNA of Cheddar Man was discovered to be haplogroup U5b1 by a Natural History Museum study in 2018 using next generation sequencing. Some 65% of western European Mesolithic hunter-gatherers had haplogroup U5; today it is widely distributed, at lower frequencies, across western Eurasia and northern Africa. In 1996, Bryan Sykes of the University of Oxford first sequenced the mitochondrial DNA from one of Cheddar Man's molars as U5a using PCR testing. The difference between the older result and the 2018 Natural History Museum result was attributed to the use of older PCR technology and possible contamination. In popular culture Soon after the discovery of the skeleton, Cheddar Man was heralded as the 'first Englishman' with an initially proposed age of 40,000-80,000 years, and became part of a discourse of British (or English) nationalism and cultural heritage. The analysis of Cheddar Man's mitochondrial DNA by Bryan Sykes in 1996 was broadcast on a regional television programme in the UK, Once Upon a Time in the West. The programme emphasised the connection between Cheddar Man and a history teacher from a local school, both of whom