of Villanovan culture date back to circa 900 BCE. In the region south of the Tiber (Latium Vetus), the Latial culture of the Latins emerged, while in the north-east of the peninsula the Este culture of the Veneti appeared. Roughly in the same period, from their core area in central Italy (modern-day Umbria and Sabina region), the Osco-Umbrians began to emigrate in various waves, through the process of Ver sacrum, the ritualized extension of colonies, in southern Latium, Molise and the whole southern half of the peninsula, replacing the previous tribes, such as the Opici and the Oenotrians. This corresponds with the emergence of the Terni culture, which had strong similarities with the Celtic cultures of Hallstatt and La Tène. The Umbrian necropolis of Terni, which dates back to the 10th century BCE, was identical in every aspect to the Celtic necropolis of the Golasecca culture. Antiquity By the mid-first millennium BCE, the Latins of Rome were growing in power and influence. This led to the establishment of ancient Roman civilization. In order to combat the non-Italic Etruscans, several Italic tribes united in the Latin League. After the Latins had liberated themselves from Etruscan rule they acquired a dominant position among the Italic tribes. Frequent conflict between various Italic tribes followed. The best documented of these are the wars between the Latins and the Samnites. The Latins eventually succeeded in unifying the Italic elements in the country. Many non-Latin Italic tribes adopted Latin culture and acquired Roman citizenship. During this time Italic colonies were established throughout the country, and non-Italic elements eventually adopted the Latin language and culture in a process known as Romanization. In the early first century BCE, several Italic tribes, in particular the Marsi and the Samnites, rebelled against Roman rule. This conflict is called the Social War. After Roman victory was secured, all peoples in Italy, except for the Celts of the Po Valley, were granted Roman citizenship. In the subsequent centuries, Italic tribes were assimilated into Latin culture in a process known as Romanization. Genetics A genetic study published in Science in November 2019 examined the remains of six Latin males buried near Rome between 900 BC and 200 BC. They carried the paternal haplogroups R-M269, R-311, R-PF7589 and R-P312 and the maternal haplogroups H1aj1a, T2c1f, H2a, U4a1a, H11a and H10. A female from the preceding Proto-Villanovan culture carried the maternal haplogroups U5a2b. These examined