penthouse in the Bogon Galaxy, Ratchet (James Arnold Taylor) and Clank (David Kaye) receive word that Ratchet's home planet, Veldin, is under attack by an army of Tyhrranoids. Posing as an officer in the Galactic Rangers, he leads an assault that drives off the invaders, who appear to use very sophisticated weapons and equipment. At a post-victory briefing, the Galactic President informs the duo that a robotic villain known as Dr. Nefarious (Armin Shimerman) is believed to be responsible for organizing the invasion. As very little is known about Nefarious, the Rangers decide to track down the only man to have ever beaten him: Captain Qwark (Jim Ward). Ratchet discovers a feral Qwark living with an indigenous tribe on planet Florana and takes him to the Starship Phoenix, where he meets Ranger Captain, Sasha Phyronix (Leslie Carrara-Rudolph). Upon regaining his sanity after watching his old adventures reenacted via vid-comics, Qwark forms the "Q-Force", an "elite" squad that answers only to him, and orders Ratchet, Clank, and Q-Force operative Skidd McMarx (Neil Flynn) to infiltrate Nefarious's base on planet Aquatos. Hacking his database, Clank obtains a map of Tyhrranosis, the Tyhrranoid home world. After Ratchet succeeds in killing the Tyhrranoid queen, Dr. Nefarious sends a transmission, mocking the Q-Force; Big Al (Chris Hatfield), the technical expert of the team, traces it to a munitions factory on planet Daxx. There, Ratchet and Clank learn that Nefarious has constructed a weapon called the Biobliterator, but they are unable to find any information on it. They also find a music video produced by pop star Courtney Gears (Melissa Disney) (a robotic parody of Britney Spears), in which she instructs robots to destroy organic lifeforms. Clank sets up a meeting with Gears, but she renders him unconscious and turns him over to Dr. Nefarious. Aware of Clank's fame as the star of "Secret Agent Clank", Nefarious asks him to serve as the face of his plan to wipe out all organic races, but he refuses. In response, Nefarious creates an evil double named Klunk (also voiced by Kaye) and has him take Clank's place. Unaware of the deception, Ratchet undertakes a mission with Skidd to investigate enemy activity on the Obani Moons. When Ratchet returns to the Phoenix, Gears abducts Skidd and turns him into a robot. Ratchet confronts and disables her in combat, then gets Skidd to safety. Meanwhile, Sasha locates Nefarious' ship, the