increasingly prominent over time. Mut, too, could appear in both leonine and cat form. Likewise, cobra goddesses often represented the Eye of Ra. Among them was Wadjet, a tutelary deity of Lower Egypt who was closely associated with royal crowns and the protection of the king. Other cobra goddesses associated with the eye include the fertility deity Renenutet, the magician goddess Weret-hekau, and Meretseger, the divine protector of the burial grounds near the city of Thebes. The deities associated with the eye were not restricted to feline and serpent forms. Hathor's usual animal form is a cow, as is that of the closely linked eye goddess Mehet-Weret. Nekhbet, a vulture goddess, was closely connected with Wadjet, the eye, and the crowns of Egypt. Many eye goddesses appear mainly in human form, including Neith, a sometimes warlike deity sometimes said to be the mother of the sun god, and Satet and Anuket, who were linked with the Nile cataracts and the inundation. Other such goddesses include Sothis, the deified form of the star of the same name, and Maat, the personification of cosmic order, who was connected with the eye because she was said to be the daughter of Ra. Even Isis, who is usually the companion of Osiris rather than Ra, or Astarte, a deity of fertility and warfare who was imported from Canaan rather than native to Egypt, could be equated with the solar eye. Frequently, two eye-related goddesses appear together, representing different aspects of the eye. The juxtaposed deities often stand for the procreative and aggressive sides of the eye's character, as Hathor and Sekhmet sometimes do. Wadjet and Nekhbet can stand for Lower and Upper Egypt, respectively, along with the Red Crown and White Crown that represent the two lands. Similarly, Mut, whose main cult center was in Thebes, sometimes served as an Upper Egyptian counterpart of Sekhmet, who was worshipped in Memphis in Lower Egypt. These goddesses and their iconographies frequently mingled. Many combinations such as Hathor-Tefnut, Mut-Sekhmet, and Bastet-Sothis appear in Egyptian texts. Wadjet could sometimes be depicted with a lion head rather than that of a cobra, Nekhbet could take on cobra form as a counterpart of Wadjet, and a great many of these goddesses wore the sun disk on their heads, sometimes with the addition of a uraeus or the cow horns from Hathor's typical headdress. Beginning in the Middle Kingdom, the hieroglyph