vary in size over the body. Conical limpet-like scales are only preserved on a fold of skin preserved on the back of the tibia, but which was probably from the bottom of the belly, rather than the leg. Separating the polygonal scales of C. casuarius are shieldlike scales, arranged close together in rows. Ossified tendons are present on all the vertebrae, except for those in the cervical region. On no vertebrae do the tendons extend below the transverse processes. Each tendon is flattened at its origin, and transversely ovoid in the central rod, ending at a rounded point. Aside from those found on Corythosaurus casuarius, extensive skin impressions have been found on Edmontosaurus annectens and notable integument has also been found on Brachylophosaurus canadensis, Gryposaurus notabilis, Parasaurolophus walkeri, Lambeosaurus magnicristatus, L. lambei, Saurolophus angustirsotris and on unidentified ornithopods. Of these, L. lambei, C. casuarius, G. notabilis, P. walkeri, and S. angustirsotris have preserved polygonal scales. The scales on L. lambei, S. angustirostris and C. casuarius are all similar. Corythosaurus is one of very few hadrosaurids which have preserved skin impressions on the hind limbs and feet. A study in 2013 showed that amongst hadrosaurids, Saurolophus angustirostris preserved the best and most complete foot and limb integument, although other species like S. osborni, Edmontosaurus annectens and Lambeosaurus lambei (= L. clavinitialis) share a fair amount of preserved tissue on those regions. It was once thought that this dinosaur lived mostly in the water, due to the appearance of webbed hands and feet. However, it was later discovered that the so-called "webs" were in fact deflated padding, much like that found on many modern mammals. Distinguishing characteristics A set of characters were indicated by Barnum Brown in 1914 to distinguish Corythosaurus from all other hadrosaurids from Alberta: a comparatively short skull with a high helmet-like crest formed by the nasals, prefrontals and frontals; the nasals not being separated in front by the premaxillaries; a narrow beak, with an expansion in front of an elongated naris; and a small narial opening. In 1916, Brown expanded the character set to include more features; in the revised version: a comparatively short skull with a high helmet-like crest formed by nasals, prefrontals and frontals; the nasals not being separated in front by premaxillaries; a narrow beak, expanded section in front of the elongated nares; a small narial opening; a vertebral formula of 15 cervicals, 19 dorsals,