component (particularly the distinctive denticulated microliths), and the later phases by the presence of uniserial (phase5) and biserial 'harpoons' (phase6) made of bone, antler and ivory. Debate continues about the nature of the earliest Magdalenian assemblages, and it remains questionable whether the Badegoulian culture is the earliest phase of Magdalenian culture. Similarly, finds from the forest of Beauregard near Paris have been suggested as belonging to the earliest Magdalenian. The earliest Magdalenian sites are in France. The Epigravettian is a similar culture appearing at the same time. Its known range extends from southeast France to the western shores of the Volga River, Russia, with many sites in Italy. The later phases of Magdalenian culture are contemporaneous with the human re-settlement of north-western Europe after the Last Glacial Maximum during the Late Glacial Maximum. As hunter gatherers, Magdalenians did not re-settle permanently in northwest Europe, instead following herds and seasons. By the end of the Magdalenian epoch, lithic technology shows a pronounced trend toward increased microlithisation. The bone harpoons and points have the most distinctive chronological markers within the typological sequence. As well as flint tools, Magdalenians are known for their elaborate worked bone, antler and ivory that served both functional and aesthetic purposes, including perforated batons. The sea shells and fossils found in Magdalenian sites may be sourced to relatively precise areas and have been used to support hypotheses of Magdalenian hunter-gatherer seasonal ranges, and perhaps trade routes. In northern Spain and south-west France this tool culture was superseded by the Azilian culture. In northern Europe it was followed by variants of the Tjongerian techno-complex. It has been suggested that key Late-glacial sites in south-western Britain may be attributed to Magdalenian culture, including Kent's Cavern. Art Bones, reindeer antlers and animal teeth display crude pictures carved or etched on them of seals, fish, reindeer, mammoths and other creatures. The best of Magdalenian artworks are a mammoth engraved on a fragment of its own ivory; a dagger of reindeer antler, with a handle in form of a reindeer; a cave-bear cut on a flat piece of schist; a seal on a bear's tooth; a fish drawn on a reindeer antler; and a complete picture, also on reindeer antler, showing horses, an aurochs, trees, and a snake biting a man's leg. The man is naked, which, together with the snake, suggests a warm climate in spite of the presence of the reindeer.