taken by historians as symbolic of the loyalty that Alan Rufus displayed to William, and from this time Richmond would remain in the hands of the most loyal of English kings' nobles and also represent a means for the King to allocate wealth to his closest "kin-group" in the sense defined by Keats-Rohan. Alan Rufus emerged as the wealthiest and most important of the Breton nobles around William I, was a key supporter of William II in the events preceding, during and following the Rebellion of 1088 and may have participated in the invasion of Normandy in 1091. Alan Rufus died on 4 August 1093 due to an unknown cause. His succession settled quickly upon his younger brother, another Alan, nicknamed "Niger" (The Black), who seems to have died by 1098. Stephen, their younger brother, inherited Richmond. Stephen died between 1135 and 1138, and was succeeded in Brittany by his eldest son, Geoffrey Boterel II, a supporter of the Empress Matilda, and in England by a younger son, Alan, also nicknamed The Black, who was an ally of King Stephen during The Anarchy. The Penthièvre brothers who held the territorial designation as Lords of Richmond are often reckoned as de facto 'earls of Richmond', though they were not so in the later strictly legal sense. Through the reign of King Henry I few earldoms had been created. King Stephen's reign was marked by the creation of several new earldoms. Stephen of Tréguier's son Alan (c. 1116–1146), was the first of these lords to be styled "Earl of Richmond" in a strictly legal sense. King Stephen also created Alan 1st Earl of Cornwall although this title would be forfeit in 1141 after the Battle of Lincoln. Alan of Brittany is recorded as riding at King Stephen's side in the Battle of Lincoln. Alan Rufus, Stephen of Tréguier, and Alan were members of "the Richmond-Penthievre... [family, and this family]... maintained its rivalry with its ducal cousins [in Brittany] into the next century". Alan married Bertha, daughter and heiress of Duke Conan III of Brittany. Alan died in 1146 at which time his widow returned to Brittany. Their son, Duke Conan IV of Brittany (c. 1138–1171) married Margaret of Huntingdon, sister of King Malcolm IV of Scotland. Conan asserted his right to Brittany, and with it Richmond; he transferred the title Earl of Richmond during his lifetime to his daughter Constance (c. 1161–1201).