2015 study by Reeder et al.: it closely integrated morphological, molecular, and paleontological data in a large dataset to overcome previous conflicts, which revealed new morphological support for molecular results that recovered Mosasauria as a sister clade to Serpentes. Another approach was developed by R. Alexander Pyron in a 2016 study, which also recovered Mosasauria as a sister clade to the Serpentes. Lower classifications The traditional view of mosasaur evolution held that all paddle-limbed (hydropedal) mosasaurs originated from a single common ancestor with functional legs (plesiopedal). However, this was shaken with the discovery of Dallasaurus, a plesiopedal mosasauroid that was found by multiple phylogenetic analyses to be an ancestor of the Mosasaurinae but unrelated to other mosasaurs. Bell and Polycn (2005) grouped these outside mosasaurs into two clades: the Russellosaurina, whose basal members include plesiopedal genera (Tethysaurinae) of their own and derived members consisting of the Plioplatecarpinae and Tylosaurinae; and the Halisauromorpha, containing the Halisaurinae. The placement of Dallasaurus suggested that the Russellosaurina and Halisauromorpha may have evolved a hydropedal form independently, the former through the tethysaurines, meaning that their placement within the Mosasauridae creates an unnatural polyphyly and thus potentially invalid. Caldwell informally proposed in a 2012 publication that the definition of a mosasaur must thus be redefined into one that does not consider russellosaurines and halisauromorphs as true mosasaurs, but as an independent group of marine lizards. However, phylogenetic studies of mosasaurs can be fickle, especially when wild card taxa like Dallasaurus remain poorly understood. For example, some studies such as a 2009 analysis by Dutchak and Caldwell instead found that Dallasaurus was ancestral to both russellosaurines and mosasaurines, although results were inconsistent in later studies. A 2017 study by Simoes et al. noted that utilization of different methods of phylogenetic analyses can yield different findings and ultimately found a consistent indication that tethysaurines were a case of hydropedal mosasaurs reversing back to a plesiopedal condition rather than an independent ancestral feature. Phylogeny Cladogram of the Mosasauroidea from Madzia and Cau (2017): Distribution Though no individual genus or subfamily is found worldwide, the Mosasauridae as a whole achieved global distribution during the Late Cretaceous with many locations typically having complex mosasaur faunas with multiple different genera and species in different ecological niches. Two African countries are particularly rich in mosasaurs: Morocco and Angola. In contemporary culture Mosasaurs, along with other extinct aquatic Mesozoic reptiles, are often erroneously depicted