Herodotus notes that Cyrus also subdued and incorporated Sogdia into the empire during his military campaigns of 546–539 BC. With Astyages out of power, all of his vassals (including many of Cyrus's relatives) were now under his command. His uncle Arsames, who had been the king of the city-state of Parsa under the Medes, therefore would have had to give up his throne. However, this transfer of power within the family seems to have been smooth, and it is likely that Arsames was still the nominal governor of Parsa under Cyrus's authority—more a Prince or a Grand Duke than a King. His son, Hystaspes, who was also Cyrus's second cousin, was then made satrap of Parthia and Phrygia. Cyrus the Great thus united the twin Achaemenid kingdoms of Parsa and Anshan into Persia proper. Arsames lived to see his grandson become Darius the Great, Shahanshah of Persia, after the deaths of both of Cyrus's sons. Cyrus's conquest of Media was merely the start of his wars. Lydian Empire and Asia Minor The exact dates of the Lydian conquest are unknown, but it must have taken place between Cyrus's overthrow of the Median kingdom (550 BC) and his conquest of Babylon (539 BC). It was common in the past to give 547 BC as the year of the conquest due to some interpretations of the Nabonidus Chronicle, but this position is currently not much held. The Lydians first attacked the Achaemenid Empire's city of Pteria in Cappadocia. Croesus besieged and captured the city enslaving its inhabitants. Meanwhile, the Persians invited the citizens of Ionia who were part of the Lydian kingdom to revolt against their ruler. The offer was rebuffed, and thus Cyrus levied an army and marched against the Lydians, increasing his numbers while passing through nations in his way. The Battle of Pteria was effectively a stalemate, with both sides suffering heavy casualties by nightfall. Croesus retreated to Sardis the following morning. While in Sardis, Croesus sent out requests for his allies to send aid to Lydia. However, near the end of the winter, before the allies could unite, Cyrus the Great pushed the war into Lydian territory and besieged Croesus in his capital, Sardis. Shortly before the final Battle of Thymbra between the two rulers, Harpagus advised Cyrus the Great to place his dromedaries in front of his warriors; the Lydian horses, not used to the dromedaries' smell,