her "the first mouse to ever get a cat in the history of the rat's kingdom". Billy, the leader of the rats. He is initially very strict about fear, calling Penny a coward for failing to gnaw the bridge, but when Penny proves her courage by inadvertently dropping a pot on Marcus's cat, he becomes extremely appreciative of her. He is voiced by Eddie Frierson in the English version. Paul, the leader of the squirrels. He is very competitive against the rats. He offers to listen to every idea suggested, and he is a reliable ally to the animals. He is voiced by Mathew Stone in the English version. Mary, an owl wearing a locket, is obsessed with her appearance and beauty. At times she is arrogant, especially to Jacob, whom she sometimes teases, but in the long run, she is sweet and sympathetic. She is voiced by Eiko Masuyama in the Japanese version and by Lisa Paulette in the English version. Carl, a green rabbit. He participates in the forest plot alongside his grandfather, and he is obedient and loyal both to him and to Peter. He is voiced by Mami Koyama in the Japanese version and by Wendee Lee in the English version. His grandfather is voiced by Richard Barnes. Jay, as his name suggests, is a blue jay. He is easily excited and upset, which is his biggest flaw, and sometimes, he cannot tell how the other animals feel. He is voiced by Stephen Apostolina in the English version. Stanley, a hedgehog. When he hears an idea or has a particular goal, he will hold onto it until it is done. His obsessive nature, however, can lead him to occasional temper tantrums. He is voiced by Don Warner in the English version. Adam, a frog. He is relatively calm, unlike Jay. When he hops, his speech is sometimes fragmented. He is voiced by Dave Mallow in the English version. Humans Marcus, the leader of the villainous humans. His leadership and courage match those of Jacob, but unlike him, Marcus is arrogant and usually doesn't think about the consequences first. He is voiced by Cyn Branch. His wife, Bertha, is voiced by Lisa Paulette. Timothy, a businessman. Like Marcus, he is intent on receiving money, and he will often ignore the consequences of his actions. He is voiced by Steve Kramer. His wife is voiced by Lara Cody.