out from what stratigraphic section the specimen (or any other fossil from Koro Toro) was first deposited in, in order to accurately radiometrically date it. Nonetheless, Abel was redated in 2010 using the same methods to about 3.65 million years ago, and Brunet agreed with an age of roughly 3.5 million years ago. A. bahrelghazali was the first australopithecine recovered from Central Africa, and disproved the earlier notion that they were restricted to east of the eastern branch of the East African Rift which formed in the Late Miocene. Koro Toro is situated about from the Rift Valley, and the remains suggest australopithecines were widely distributed in grassland and woodland zones across the continent. The lack of other Central and West African australopithecines may be due to sampling bias, as similarly aged fossil-bearing sediments are more or less unknown beyond East Africa. The ancestors of A. bahrelghazali may have left East Africa via the Central African Shear Zone. In 2014, the first australopithecine in the western branch of the East African Rift was reported in Ishango, Democratic Republic of the Congo. At present, the classification of Australopithecus and Paranthropus species is in disarray. Australopithecus is considered a grade taxon whose members are united by their similar physiology rather than how close they are to each other in the hominin family tree. In an attempt to resolve this, in 2003, Spanish writer Camilo José Cela Conde and evolutionary biologist Francisco J. Ayala proposed splitting off the genus "Praeanthropus" and including A. bahrelghazali alongside Sahelanthropus (the only other fossil ape known from Chad), A. anamensis, A. afarensis, and A. garhi. The validity of A. bahrelghazali has not been widely accepted given how few remains there are and how similar they are to A. afarensis. Anatomy The teeth of KT12/H1 are quite similar to the jawbone of A. afarensis, with large and incisor-like canines and bicuspid premolars (as opposed to molar-like premolars). Unlike A. afarensis, the alveolar part of the jawbone where the tooth sockets are is almost vertical as opposed to oblique, possesses poorly developed superior transverse torus and moderate inferior torus (two ridges on the midline of the jaw on the tongue side), and thin enamel on the chewing surface of the premolars. Brunet and colleagues had listed the presence of 3 distinct tooth roots as a distinguishing characteristic, but the third premolar of the A. afarensis LH-24 specimen from Middle