Perenniporia), Polyporus, and Fistulina. However, all of the original genera except Boletus have since been transferred to different families, and several new Boletaceae genera have been described. Genera Rolf Singer, in the 4th edition (1986) of his Agaricales in Modern Taxonomy, included 26 genera and 415 species in Boletaceae. In the Dictionary of the Fungi (10th edition, 2008), 35 Boletaceae genera were recognised, which collectively contained 787 species. Molecular phylogenetic studies in the 2000s have revised the concept of the family; in a highly cited 2006 publication, Manfred Binder and David Hibbett recognised 38 genera within the family, many of which had remained at the time undescribed. The number of Boletaceae genera increased significantly in the following years, as some of the early-established genera (Boletus, Leccinum, Xerocomus), further revealed to be highly polyphyletic. In the comprehensive work of Wu and colleagues (2014), seven major clades at subfamily level and 59 generic lineages were uncovered, including four new subfamilies (Austroboletoideae, Chalciporoideae, Leccinoideae, and Zangioideae) and 22 potential new genera. To formally name the generic lineages unravelled by molecular phylogenies, several new genera have since been described from Asia, Europe and North America including, among others, Baorangia, Butyriboletus, Cacaoporus, Caloboletus, Exsudoporus, Imperator and Rubroboletus. Some characters traditionally emphasised in morphology-based taxonomy, such as basidiospore ornamentation and "stuffed" pore morphology, revealed to be incongruent with molecular taxonomy, suggesting that certain traits evolved more than once within the family. (*) Note that the phylogenetic and taxonomic position of many taxa currently remaining in genus Boletus has not yet been clarified. The number of species in this genus will therefore significantly reduce in the following years, as more taxa will be transferred to different genera, or found to be synonyms. Many other genera formerly part of this family have been moved into other, smaller families, as work with molecular phylogeny shows that they are more distantly related, even if morphologically similar. Representative of this adjustment, is the move of the slimy-capped genus Suillus to the family Suillaceae. Distribution Boletes are found worldwide, on every continent except Antarctica. Well-known and well-described in the temperate latitudes in the northern hemisphere, newer research has shown significant diversity in tropical and southern hemisphere regions as well. E. J. H. Corner found evidence of at least 60 species on the island of Singapore alone. In 1972 he described 140 species from the Malay Peninsula and Borneo and estimated there were