the specific name. Pairs eight, nine and ten again equalled the first. After a narrow hiatus between the second and third snout fragment four tooth sockets are present at each side of the latter, but these are not placed in opposite pairs. The number of teeth in the upper jaw thus seems to have been at least fourteen. The smaller sized teeth were placed in a constriction of the snout, which thus had a broader end with larger teeth, a so-called "prey grab", usually interpreted as an adaptation to catch slippery prey such as fish. The posterior skull fragment, a braincase which is rather damaged, shows on its top the base of a parietal crest, probably pointing towards the back. It seems to have been separate from the snout crest. The layer the fossils were found in, does not consist of marine sediments, but contains land plant debris; this is seen as an indication of a more terrestrial habitat. David Martill estimated Caulkicephalus had a wingspan of around 5 meters (16.5 ft). Classification Caulkicephalus was by the describers assigned to the Ornithocheiridae in view of the narrowing in the middle of the snout. The snout crest was seen as an indication it belonged to the more general Ornithocheiroidea sensu Unwin, whereas the parietal crest was suggested to have been a synapomorphy, a shared new feature, of the more narrow group of the Euornithocheira. Unique characters of the species itself, its autapomorphies, are the details of its dentition, the downward and backward running suture between the premaxilla and maxilla, and the fact the median ridge of the palate begins (or ends) at the ninth tooth pair. In 2019 however, several studies have reassigned Caulkicephalus to the family Anhangueridae, specifically to the subfamily Anhanguerinae, sister taxon to both Guidraco and Ludodactylus. The cladogram below is a topology recovered by Rodrigo Pêgas and colleagues in 2019, showing the position of Caulkicephalus within the Anhangueridae. See also List of pterosaur genera Timeline of pterosaur research Pterosaur size References External links Description, references, and images of Caulkicephalus. Pteranodontoids Early Cretaceous pterosaurs of Europe Fossil taxa described in 2005