comprise up to 22 individuals. The gaur (Bos gaurus) is the largest ungulate in the park, with herds of up to 42 individuals that frequent foremost grasslands in the vicinity of water sources. The sambar deer (Cervus unicolor) forms smaller groups of up to five individuals, but also congregates in groups of up to 45 individuals in the wet season. The chital (Axis axis) forms large groups of at least 35 individuals, with some herds increasing to more than 100 members in the wet season. Chital, Indian spotted chevrotain (Moschiola indica) and Indian muntjac (Muntiacus muntjak) have been recorded eating fallen fruit of the Indian gooseberry in a forest monitoring plot; they are therefore considered to be the primary seed dispersers in the park. Present are also four-horned antelope (Tetracerus quadricornis), blackbuck (Antilope cervicapra), wild boar (Sus scrofa), Indian pangolin (Manis crassicaudata) and Indian crested porcupine (Hystrix indica). Four bonnet macaque (Macaca radiata) troops were studied in 1997, which ranged in size from 28 to 35 members and lived in sympatry with gray langur (Semnopithecus entellus) troops. A troop in the Moyar River valley foraged on leaves, flowers and fruits of several tree and shrub species including tamarind (Tamarindus indica), banyan fig (Ficus benghalensis), wild jujube (Ziziphus oenoplia), neem (Azadirachta indica), kaayam (Memecylon edule) and indigoberry (Randia malabarica), but also consumed herbs, crickets and grasshoppers. The range of the Indian giant squirrel (Ratufa indica) is continuous in the national park's moist deciduous forest; in the drier eastern part, it inhabits foremost riverine habitat with contiguous canopy. It builds nests in trees with a mean canopy height of and feeds on 25 plant species including teak, Indian laurel and Grewia tiliifolia. The Indian giant flying squirrel (Petaurista philippensis) inhabits foremost moist deciduous forest with old trees of a mean height, a mean density of and a canopy height of at least . In 2013, a painted bat (Kerivoula picta) was sighted in the eastern part of the tiger reserve. Birds Birds observed from 1994 to 1996 comprised 266 species; the 213 resident ones include Malabar grey hornbill (Ocyceros griseus), Indian grey hornbill (O. birostris), Indian peafowl (Pavo cristatus), Bonelli's eagle (Aquila fasciata), crested serpent eagle (Spilornis cheela), black eagle (Ictinaetus malaiensis), besra (Accipiter virgatus) and crested goshawk (A. trivirgatus), white-rumped shama (Copsychus malabaricus), Indian roller (Coracias benghalensis), greater flameback (Chrysocolaptes guttacristatus) and white-naped woodpecker (C. festivus), black-rumped flameback (Dinopium benghalense), white-bellied