Megaladapis, informally known as the koala lemur, was a genus belonging to the family Megaladapidae, consisting of three extinct species of lemurs that once inhabited the island of Madagascar. The largest measured between in length. Adaptations Megaladapis was quite different from any living lemur. Its body was squat and built like that of the modern koala. Its long arms, fingers, feet, and toes were specialized for grasping trees, and its legs were splayed for vertical climbing. The hands and feet were curved and the ankles and wrists did not have the usual stability needed to travel on the ground that most other lemurids have. Its pedal morphology suggests Megaladapis evolved to live in an arboreal environment. Its foot had a large hallux and lateral abductor musculature that helped it to grasp vertically on trees, features shared by other arboreal species. Its head was unlike that of any other primate; most strikingly, its eyes were on the sides of its skull, instead of forward on the skull as in all other primates. Its long canine teeth and cow-like jaw formed a tapering snout. Its jaw muscles were powerful for chewing the tough native vegetation. Based on the microwear patterns of its teeth, Megaladapis is believed to have been folivorous, using a leaf-cropping foraging method. These patterns found no permanent upper incisors or the presence of an expanded articular facet on the posterior face of the mandibular condyle. This diet and similar phenotypic traits of the teeth are the basis for concluding a shared ancestry with the Lepilemur. The diet, however, might be the factor that influences the dental development. Species with a larger brain, later initiation of molar crowns, and longer formation of crown are considered to have more of an omnivorous diet. In contrast, Megaladapis lived on a folivorous diet, despite having a smaller brain, early initiation of molar crowns, and fast crown formation. Its body weight reached . Other estimates suggest but its still much larger than any extanct lemur. The shape of its skull was unique among all known primates, with a nasal region which showed similarities to those of rhinoceros, a feature that probably combined with an enlarged upper lip for grasping leaves. It had the largest body size of any lemur, with double the body mass of the next largest extinct lemur. An endocast of its skull showed that it had a brain capacity of about