The Syracusan fleet positioned in Issa was informed in time, and Greek triremes attacked the siege fleet, taking victory in the end. According to Diodorus, the Greeks killed more than 5,000 and captured 2,000 prisoners, ran down or captured their ships, and burned their weapons in dedication to their god. This battle meant the loss of the most important strategic Liburnian positions in the centre of the Adriatic, resulting in their final retreat to their main ethnic region, Liburnia, and their complete departure from the Italic coast, apart from Truentum (nowadays on the border between Marche and Abruzzo). Greek colonization, however, did not extend into Liburnia, which remained strongly held, and Syracusan dominance suddenly diminished upon the death of Dionysius the Elder. The Liburnians recovered and developed piracy to secure navigable routes in the Adriatic, as recorded by Livius for 302 BC. The middle of the 3rd century BC was marked by the rise of an Illyrian kingdom in the south of the Adriatic, led by king Agron of the Ardiaei. Its piratical activities imperiled Greek and Roman interests in the Adriatic, and caused the first Roman intervention on the eastern coast in 229 BC; Florus in Epitome of Roman History noted the Liburnians as the Romans' enemies in this expedition, while Appian (Bell. Civ., II, 39) noted liburnae as swift galleys the Romans first fought with when they entered the Adriatic. The Liburni were allies of their southern Illyrian compatriots, Ardiaei and the others, but from the lack of more records related to them in the 3rd century BC, it is assumed that they mostly stood aside in the subsequent Roman wars and conflicts with Pyrrhus, Carthage, Macedonia and the southern Illyrian state. Even though Liburnian territory was not involved in these confrontations, it seems that the Liburna warship was adopted by the Romans during the Punic Wars and in the Second Macedonian War. Hellenistic and Roman periods In 181 BC, the Romans established their colony at Aquileia and took control of all Venetia in the north, thus expanding towards the Illyrian area from the northwest. In 177 BC they conquered Istria to the north of the eastern Adriatic coast, settled by tribe of Histri, while the Iapodes, the northern neighbors of Liburnia, attacked Aquileia in 171 BC. These incidents did not involve Liburnian territory. The Liburnians probably avoided direct conflict with the Romans in order to safeguard their