is currently classified as Endangered by the IUCN. The species is now absent from many parts of its former range, particularly the savanna country. In 2005 it was listed as Endangered under Australian Commonwealth legislation (EPBC Act). Threats are predation by feral cats, dingoes and foxes, particularly after fire or grazing has removed protective ground cover. Destruction, degradation, and fragmentation of the quoll's habitat due to land clearing, grazing, pasture improvement, and mining are also significant. Quolls are also susceptible to being run over on roads. The current major threat to the northern quoll in the northern and western portion of its range is the spread of cane toads. Like many other native Australian species, northern quolls are poisoned after eating or mouthing cane toads. Cane toads were originally introduced in Queensland, but have now occupied the Top End of the Northern Territory, including Kakadu National Park and the Darwin area, and entered the Kimberley region of Western Australia, where they are established around Kununurra and Lake Argyle. Immediately after cane toad invasion of Kakadu, quolls became extinct at one study site and declined from 45 individuals to five at another site. The northern quoll may cease to exist in most areas in the Top End once the cane toad population completely overlaps the northern quoll's range. However, remnant populations of northern quolls still persist in Queensland where cane toads have been present for decades. These persisting Queensland quoll populations are naturally toad averse (as observed on remote cameras). One of the northern quoll populations studied in Kakadu during the recent cane toad invasion, had a few individuals survive the invasion. These individuals were, likely similar to the Queensland quolls, genetically averse to the toads. In 2003, to help protect northern quolls, numerous quolls were transferred to the toad-free English Company Islands (Astell and Pobassoo Islands), off the coast of Arnhem Land. Quolls thrived on these islands, with an estimated population of 2193 female northern quolls on Astell Island by 2014. In 2017, quolls from Astell Island were collected, trained via conditioned taste aversion to avoid attacking cane toads and reintroduced to Kakadu National Park. This reintroduction attempt saw the release both toad-trained (22) and toad-naive (9) northern quolls to an area of Kakadu that previous had quolls but where they had recently gone locally extinct due to the invasion of cane toads. Although the toad-trained quolls survived longer than