ocean. In Washington and Oregon, they commonly nest in Douglas-fir (Pseudotsuga menziesii) dominated stands. They also select stands dominated by mountain hemlock (Tsuga mertensiana), western redcedar (Thuja plicata), and Sitka spruce (Picea sitchensis) for nesting. In California, nests are most often located in redwood (Sequoia sempervirens) dominated stands with scattered Sitka spruce, western hemlock (Tsuga heterophylla) and Douglas-fir. Marbled murrelets also occur in stands dominated by Port-Orford-cedar (Chamaecyparis lawsoniana). Foraging habitat Marbled murrelets forage in the ocean near shore and in inland saltwater areas such as bays, sounds, and saltwater passageways. Some also forage on inland freshwater lakes. Flocks of 50 or more birds have been observed near freshwater lakes. Subadults occur at sea throughout the summer. Marbled murrelets feed within 1,640 feet (500 m) of shore. Winter habitat Marbled murrelet winter habitat is the same as the nesting and foraging habitat. During the winter marbled murrelets use inland old-growth or mature sites for roosting, courtship, and investigating nest sites. The use of inland lakes during the nonbreeding season occurs in conjunction with visits to nesting areas. Behavior Food and feeding The marbled murrelet feeds at sea both in pelagic offshore areas (often associating with upwellings) and inshore in protected bays and fiords. The bird has not been known to wander from the Pacific coast of North America, all inland and eastern Brachyramphus records being of the closely related long-billed murrelet. They feed below the water surface on small fish and invertebrates. Some principal foods include sand lance (Ammodytes hexapterus), Pacific herring (Clupea haringus), capelin (Mallotus villosus), shiner perch, and the invertebrates Euphausia pacifica and Thysanoessa spinifera. They often forage in pairs but do not feed in large flocks as do other alcids. Loose aggregations of 500 or more birds occasionally occur in winter. Subadults feed singly; but in early July, when pairs of adults are still feeding young, mixed flocks begin to form. Marbled murrelets feed during the day and at night. Breeding The nesting behavior of the marbled murrelet is unusual, since unlike most alcids it does not nest in colonies on cliffs or in burrows, but on branches of old-growth and mature conifers such as western hemlock, Sitka spruce, Douglas-fir and coastal redwood, as far as 80 km inland. It lays one egg on a platform of lichen or moss on these branches (less often on the ground). In northern populations, murrelets nest on the ground among