ignicapilla and Mediterranean R. i. balearicus (Jordans, 1923). The latter form is found on the Balearic Islands and in north Africa, and is slightly paler below and greyer above than the nominate subspecies. Other subspecies have been claimed, including southeastern R. i. caucasicus, North African R. i. laeneni, and Crimean R. i. tauricus. The Madeira firecrest, R. madeirensis, was formerly also considered to be a subspecies of the common firecrest, but phylogenetic analysis based on the cytochrome b gene showed that the Madeiran form is distinct at the species level. Cytochrome b gene divergence between the Madeira firecrest and the European bird is 8.5%, comparable with the divergence level between other recognised Regulus species, such as the 9% between the goldcrest and the golden-crowned kinglet. The island form also differs in morphology and vocalisations. The proposed split was accepted by the Association of European Rarities Committees (AERC) in 2003, with most other authorities also splitting it later on. The flamecrest or Taiwan firecrest (Regulus goodfellowi) of Taiwan has sometimes been viewed as a race of the common firecrest; however, the flamecrest's territorial song, which resembles those of the Himalayan races of the goldcrest, and genetic data indicate that the flamecrest is closely related to the Himalayan goldcrest and only distantly to the two firecrest species. The kinglets on the Canary Islands, which were also considered to be close to firecrests, have now been shown to comprise two subspecies of goldcrest. Fossils There are a few Pleistocene (2.6 million to 12,000 years ago) records from Europe and Israel of extant Regulus species, mostly goldcrests or unidentifiable to species, but also a Spanish specimen of firecrest. A left ulna from Bulgaria was identified as belonging to a fossil species, Regulus bulgaricus, from 2.6–1.95 mya. This appears to be ancestral to the common firecrest, with the goldcrest diverging from this lineage in the Middle Pleistocene. Distribution and habitat The common firecrest breeds in lowland broadleaf forest, preferring cork oak and alder where available, otherwise beech and holly. It also uses mixed broadleaf and conifer woodland, and stands of spruce, European silver fir, cedar and pines, often with undergrowth of juniper, ivy and wild rose. In drier Mediterranean habitats it is found in conifers, evergreen oak, and mixed woodlands up to . Unlike more specialised birds such as Eurasian nuthatch and common treecreeper, both of which forage on trunks, the crests do not need