a comical, streetwise Bluestreak cleaner wrasse who works in the Whale Wash of Reef City and often concocts schemes to become rich. Jack Black as Lenny, Don Lino's youngest son, a vegetarian, and the younger brother of Frankie, who becomes good friends with Oscar and Angie. Renée Zellweger as Angie, a pink angelfish who harbors a secret crush on Oscar. Angelina Jolie as Lola, a seductive female lion fish whom Oscar develops a romantic interest in. Robert De Niro as Don Lino, a great white shark and leader of a mob consisting of criminally-inclined sharks, who wants his two sons to take over his business and run it together. Martin Scorsese as Sykes, a loan shark pufferfish who owns the Whale Wash and to whom Oscar owes five thousand clams. Ziggy Marley and Doug E. Doug as Ernie and Bernie, two jellyfish with Jamaican accents and Sykes' enforcers, who enjoy torturing Oscar with their painful stingers when he is in trouble with their boss. Michael Imperioli as Frankie, Lenny's older brother and Don Lino's more vicious son, who is embarrassed by Lenny's vegetarian views. Vincent Pastore as Luca, Don Lino's teal octopus "left-hand, right-hand man", with a tendency to state the obvious, much to Lino's frustration. Peter Falk as Don Feinberg, an elderly leopard shark and leader of a mob of criminally-inclined leopard sharks, who is a friend of Don Lino. Katie Couric as Katie Current, a female sea bass who is the local reporter of the Southside Reef in the U.S. release. At the time, Katie Couric co-hosted Today in America. In the Australian release, then local Today co-host Tracy Grimshaw dubbed the lines. Fiona Phillips of the UK's GMTV performed the voice for the British release of the film. Cristina Parodi of Italy's Verissimo provided the Italian version of the character. David P. Smith as Crazy Joe, a deranged hermit crab who is Oscar's best friend. Bobb'e J. Thompson, Kamali Minter, Emily Lyon Segan as the Shorties, three delinquent young fish and friends of Oscar who love to spray graffiti. Two of them are cowfishes and one is a Pennant coralfish. Shelley Morrison as Mrs. Sanchez, an old grumpy weeverfish. David Soren as an unnamed shrimp who fears being eaten by a shark and is an enemy to Don Lino. Sean Bishop as an unnamed green sperm whale who attends the Whale Wash. Christina Aguilera and Missy Elliott