been proposed and applied to determine which speciation mode a species underwent in the past using various approaches or combinations thereof: species-level phylogenies, range overlaps, symmetry in range sizes between sister species pairs, and species movements within geographic ranges. Molecular clock dating methods are also often employed to accurately gauge divergence times that reflect the fossil or geological record (such as with the snapping shrimp separated by the closure of the Isthmus of Panama or speciation events within the genus Cyclamen). Other techniques used today have employed measures of gene flow between populations, ecological niche modelling (such as in the case of the Myrtle and Audubon's warblers or the environmentally-mediated speciation taking place among dendrobatid frogs in Ecuador), and statistical testing of monophyletic groups. Biotechnological advances have allowed for large scale, multi-locus genome comparisons (such as with the possible allopatric speciation event that occurred between ancestral humans and chimpanzees), linking species' evolutionary history with ecology and clarifying phylogenetic patterns. References Further reading Mathematical models of reproductive isolation Biogeography Ecology Evolutionary biology Speciation