Coloborhynchus is a genus of pterodactyloid pterosaur belonging to the family Anhangueridae, though it has also been recovered as a member of the Ornithocheiridae in some studies. Coloborhynchus is known from the Lower Cretaceous of England (Valanginian age, 140 to 136 million years ago), and depending on which species are included, possibly the Albian and Cenomanian ages (113 to 93.9 million years ago) as well. Coloborhynchus was once thought to be the largest known toothed pterosaur, however, a specimen of the closely related Tropeognathus is now thought to have had a larger wingspan. History and classification Like many ornithocheiroid pterosaurs named during the 19th century, Coloborhynchus has a highly convoluted history of classification. Over the years numerous species have been assigned to it, and often, species have been shuffled between Coloborhynchus and related genera by various researchers. In 1874 Richard Owen, rejecting the creation by Harry Govier Seeley of the genus Ornithocheirus, named a species Coloborhynchus clavirostris based on holotype BMNH 1822, a partial snout from the Hastings Beds of the Wealden Group of East Sussex, England. The genus name means "maimed beak", a reference to the damaged and eroded condition of the fossil; the specific name means "key snout", referring to its form in cross-section. Owen also reclassified Ornithocheirus cuvieri and O. sedgwickii as species within the genus Coloborhycnhus, though he did not designate any of these three as the type species. Owen considered the defining trait of the genus to be the location of the front tooth pairs high on the side of the upper jaws. However, in 1913 Reginald Walter Hooley concluded that this location was an artefact of the erosion and that the genus was indistinguishable from Criorhynchus simus, the second genus and species Owen erected in 1874. Hooley also ignored Owen's reassignment of the two former Ornithocheirus species, leaving them in that genus. In 1967, Kuhn agreed with Hooley that Coloborhynchus clavirostris was a synonym of Criorhynchus simus. Furthermore, Kuhn was the first to formally designate C. clavirostris as the type species of the genus, rather than one of the Ornithocheirus species. Most later researchers followed these opinions, regarding Coloborhynchus as invalid relative to Criorhynchus. This changed in 1994 when Yuong-Nam Lee named Coloborhynchus wadleighi for a snout found in 1992 in the Albian age Paw Paw Formation Texas. The revival of the genus meant that of several related species, then assigned to other genera,