genus as "Ornithocheirus" compressirostris (=Lonchodectes). In 2013, Rodrigues and Kellner considered Coloborhynchus to be monotypic, containing only C. clavirostris, and placed most other species in other genera, or declared them nomina dubia. In 2018, Jacobs et al. named a new species of Coloborhynchus, C. fluviferox from the Ifezouane Formation of the Kem Kem Group of Morocco based on a partial snout, and also tentatively referred another specimen from the same locality to a different, unnamed species. A 2020 review of Coloborhynchinae by Holgado and Pêgas moved both C. capito and C. fluviferox to a new genus, Nicorhynchus, and also referred the unnamed Ifezouane Formation coloborhynchine to N. fluviferox. However, a 2023 review of Kem Kem pterosaurs found the traits that distinguish Nicorhynchus from Coloborhynchus to be subtle enough to justify their synonymy, stating that the material was damaged and fragmentary enough to support this. List of species and synonyms Species which have been assigned to Coloborhynchus by various scientists over the years include: C. clavirostris Owen, 1874, the type species ?C. ligabuei (Dalla Vecchia, 1993) = Cearadactylus ligabuei Dalla Vecchia, 1993 [also classified as Anhanguera ligabuei] ?C. piscator (Kellner & Tomida, 2000) = Anhanguera piscator Kellner & Tomida, 2000 [also classified as Anhanguera or C. robustus] Species assigned to Coloborhynchus in the past include: C. cuvieri (Bowerbank, 1851) = Ornithocheirus sedgwicki (Newton, 1888) = Pterodactylus cuvieri Bowerbank, 1851 [now classified as Cimoliopterus] C. sedgwicki (Owen 1859) = Ornithocheirus sedgwicki (Newton, 1888) = Pterodactylus sedgwickii Owen, 1859 [now classified as Camposipterus] C. araripensis (Wellnhofer, 1985) = Santanadactylus araripensis Wellnhofer, 1985 now seen as a nomen dubium ?C. robustus (Wellnhofer, 1987) = Tropeognathus robustus Wellnhofer, 1987 [now seen as a nomen dubium] C. wadleighi Lee, 1994 [also classified as Uktenadactylus] C. moroccensis (Mader & Kellner, 1999) = Siroccopteryx moroccensis Mader & Kellner 1999 C. piscator (Kellner & Tomida, 2000) = Anhanguera piscator Kellner & Tomida, 2000 C. spielbergi Veldmeijer, 2003 [now classified as Maaradactylus] C. capito (Seeley, 1870) = Ornithocheirus capito Seeley, 1870 = "Ptenodactylus capito" Seeley, 1869 [now classified as Nicorhynchus] C. fluviferox Jacobs, Martill, Ibrahim & Longrich, 2018 [now classified as Nicorhynchus] Description The type specimen of Coloborhynchus is known only from a partial upper jaw. Therefore, according to Rodrigues and Kellner's 2008 re-evaluation on Coloborhynchus clavirostris, it can only be differentiated from its relatives based on its unique combination of tooth socket positions. In Coloborhynchus, the two front teeth