the scientific literature in 1971, by Léonard Ginsburg and Guy Mennessier. In 1974, French paleontologist Jacques Fabre based on this specimen, MNHN CNJ-71, named a new species Gallodactylus canjuersensis. The generic name combines a reference to Gallia with Greek daktylos, "finger". He concluded it was the same genus as P. suevicus, but did not revive Cycnorhamphus, judging that the latter name was unavailable because of mistakes in the diagnosis by Seeley, already pointed uit by Plieninger. P. suevicus thus became Gallodactylus suevicus. In 1976, Fabre again named the species, describing it in greater detail but not mentioning the earlier publication. This confused later researchers who mistakenly assumed that 1976 was the formal naming date. In fact the 1974 paper contains a sufficient description and the species was validly named that year. In 1983, the Ghirardis sold their entire collection to the National Museum of Natural History, France. However, in 1996, Christopher Bennett pointed out that such mistakes do not invalidate a name and that therefore Cycnorhamphus has priority, making Gallodactylus canjuersensis C. canjuersensis. In 2010 and 2012, Bennett published further re-studies of the fossils, concluding that the differences between the two species could be explained by age, sex or individual variation, and formally synonymized C. canjuersensis and C. suevicus. Description Cycnorhamphus had historically been assumed to have had long jaws with teeth at the very tip, akin to those of Pterodactylus antiquus. However, recent work on a specimen nicknamed "The Painten Pelican" has revealed that the animal possesses a very unusual jaw anatomy, with peg-like teeth at the jaw tips - blunter and stouter in older individuals -, jaw curvatures behind said teeth that form angled arcs away from the biting surface, forming thus an opening, and two poorly understood soft tissue structures occupying this opening from the upper jaw, showing mineralization. The purpose of these adaptations is unknown, but they are more obvious and well developed in adult animals. It has been speculated that the jaws functioned similar to those of openbill storks, allowing the animal to hold hard invertebrates like mollusks and either crush or bisect them. Classification As illustrated below, the results of a topology are based on a phylogenetic analysis made by Longrich, Martill, and Andres in 2018. They placed the Cycnorhamphus within the clade Euctenochasmatia, more precisely within the family Gallodactylidae, sister taxon to Normannognathus. See also List of pterosaur genera Timeline of pterosaur research