similar result to the one by Andres and Myers in 2013 where Coloborhynchus is the sister taxon of Ornithocheirus. Their cladogram is shown on the left. In 2018, Jacobs et al. also recovered a similar cladogram compared to Andres and Myers (2013), where both Coloborhynchus and Siroccopteryx were assigned to the Ornithocheiridae, however, unlike the study made by Andres and Myers, they used the name Uktenadactylus wadleighi to refer to C. wadleighi in their analysis. They published their conclusion in 2019. However, many subsequent analyses made in 2019 and 2020 have recovered Coloborhynchus within the family Anhangueridae, more specifically within the subfamily Coloborhynchinae. The cladogram on the left shows a phylogenetic analysis made by Borja Holgado and Rodrigo Pêgas in 2020. Topology 1: Pentland et al. (2019). Topology 2: Holgado & Pêgas (2020). Paleobiology Similar to most anhanguerids, Coloborhynchus is traditionally thought to be an oceanic piscivore. However one study found it among the carnivorous pterosaur taxa, as opposed to several other piscivore and insectivore species. See also List of pterosaur genera Timeline of pterosaur research References The Pterosaur Database (multiple species, scroll down) Pteranodontoids Early Cretaceous pterosaurs of Europe Taxa named by Richard Owen Fossil taxa described in 1874