veins fused, thus without M1 cell Bittacomorpha Bittacomorphella Bittacomorphinae Bittacomorpha Bittacomorpha clavipes - Eastern NA to the edge of the Rocky Mountains (though not as abundant in the Plains states) Bittacomorpha occidentalis - Pacific Northwest Bittacomorphella Bittacomorphella jonesi - New England down to North Carolina, west to Minnesota and Michigan Bittacomorphella esakii - Japan Bittacomorphella fenderiana - Queen Charlotte Island down to Northern Oregon. Bittacomorphella nipponensis - Bittacomorphella pacifica - Northern California up through Oregon (possibly Washington?) Bittacomorphella sackenii - Sierra Nevadas Bittacomorphella thaiensis - Ptychopterinae Ptychoptera Ptychoptera byersi - California Ptychoptera lenis P. l. lenis - Pacific Northwest P. l. coloradensis - Colorado/Utah Ptychoptera metallica - Central Canada, Minnesota & Michigan Ptychoptera minor - California/Idaho Ptychoptera monoensis - Northern California Ptychoptera osceola - Florida Ptychoptera pendula - British Columbia to Utah & Colorado Ptychoptera quadrifasciata - Eastern North America (Syn P. rufocinctus) Ptychoptera sculleni - Pacific Northwest Ptychoptera townesi - Washington & Oregon Ptychoptera uta - Utah Fossil Subfamilies and genera Subfamily Bittacomorphinae †Probittacomorpha †Probittacomorpha brisaci - Montagne d'Andance Miocene (Turolian), France †Probittacomorpha christenseni - Fur Formation Eocene (Ypresian), Denmark; †Zhiganka †Zhiganka comitans - Batylykh Formation Early Cretaceous (Neocomian), Russia †Zhiganka longialata - Khasurty locality Early Cretaceous (Aptian), Russia †Zhiganka woolgari - Weald Clay Early Cretaceous (Hauterivian), England Subfamily. †Eoptychopterinae †Architendipes †Architendipes tshernovskiji - Dzhil Formation Early Jurassic (Hettangian), Kyrgyzstan †Doptychoptera †Doptychoptera baisica - Zaza Formation Early Cretaceous (Aptian), Russia †Eolimnobia †Eolimnobia geinitzi - Ciechocinek Formation Early Jurassic (Toarcian), Germany †Eoptychoptera †Eoptychoptera aequidistans - Dzhil Formation Early Jurassic (Hettangian), Kyrgyzstan †Eoptychoptera altaica - Ortsog Formation Middle Jurassic (Bajocian), Mongolia †Eoptychoptera ansorgei - Daohugou Formation Middle Jurassic (Callovian), China †Eoptychoptera asiatica - Karabastau Formation Late Jurassic, Kazakhstan †Eoptychoptera aucta - Karabastau Formation Late Jurassic, Kazakhstan †Eoptychoptera braziliana - Crato Formation Early Cretaceous (Aptian), Brazil †Eoptychoptera britannica - Weald Clay Early Cretaceous (Hauterivian), England †Eoptychoptera cantabrica - Spanish amber (Albian), Spain †Eoptychoptera cretacea - Zaza Formation Early Cretaceous (Aptian), Russia †Eoptychoptera elevata - Khaya Formation Late Jurassic (Tithonian), Russia †Eoptychoptera eximia - Ciechocinek Formation Early Jurassic (Toarcian), Germany †Eoptychoptera jurassica - Daohugou Formation Middle Jurassic (Callovian), China †Eoptychoptera longifurcata - Lulworth Formation Early Cretaceous (Berriasian), England †Eoptychoptera magna - Karabastau Formation Late Jurassic, Kazakhstan †Eoptychoptera maxima - Itat Formation Middle Jurassic (Bathonian), Russia †Eoptychoptera modica - Itat Formation (Middle Jurassic (Bathonian), Russia †Eoptychoptera paramaculata - Itat Formation (Middle Jurassic (Bathonian), Russia †Eoptychoptera shurabica - Sagul Formation Early Jurassic (Toarcian), Kyrgyzstan †Eoptychoptera