flush appears in April, they remove the tender blades in small clumps. Later, when grasses are higher than , they uproot entire clumps, dust them skilfully and consume the fresh leave tops, but discard the roots. When grasses are mature in autumn, they clean and consume the succulent basal portions with the roots, and discard the fibrous blades. From the bamboos, they eat seedlings, culms and lateral shoots. During the dry season from January to April, they mainly browse on both leaves and twigs preferring the fresh foliage, and consume thorn bearing shoots of acacia species without any obvious discomfort. They feed on the bark of white thorn and other flowering plants, and consume the fruits of wood apple, tamarind, kumbhi and date palm. In Nepal's Bardia National Park, elephants consume large amounts of the floodplain grass, particularly during the monsoon season. They browse more in the dry season with bark constituting a major part of their diet in the cool part of that season. During a study in a tropical moist mixed deciduous forested area of in Assam, elephants were observed to feed on about 20 species of grasses, plants and trees. Grasses such as Imperata cylindrica and Leersia hexandra constituted by far the most predominant component of their diet. The movement and habitat utilisation patterns of an elephant population were studied in southern India during 1981–83 within a study area. The vegetation types of this area encompasses dry thorn forest at , deciduous forest at , stunted evergreen forest and grassland at . Five different elephant clans, each consisting of between 50 and 200 individuals had home ranges of between and , which overlapped. They preferred habitat where water was available and food plants were palatable. During the dry months of January to April, they congregated at high densities of up to five individuals per km2 in river valleys where browse plants had a much higher protein content than the coarse tall grasses on hill slopes. With the onset of rains in May, they dispersed over a wider area at lower densities, largely into the tall grass forests, to feed on the fresh grasses, which then had a high protein value. During the second wet season from September to December, when the tall grasses became fibrous, they moved into lower elevation short grass open forests. The normal movement pattern could be upset during years of adverse environmental conditions.