Chiapas, Oaxaca and Guatemala laid the basis for the Mesoamerican cultural area. All this was facilitated by considerable regional communications in ancient Mesoamerica, especially along the Pacific coast. During this formative period distinct religious and symbolic traditions spread, as well as the development of artistic and architectural complexes. In the subsequent Preclassic period, complex urban polities began to develop among the Maya, with the rise of centers such as Aguada fénix and Calakmul in Mexico; El Mirador, and Tikal in Guatemala, and the Zapotec at Monte Albán. During this period, the first true Mesoamerican writing systems were developed in the Epi-Olmec and the Zapotec cultures. The Mesoamerican writing tradition reached its height in the Classic Maya logosyllabic script. Mesoamerica is one of only six regions of the world where writing is known to have independently developed (the others being ancient Egypt, Peru, India, Sumer, and China). In Central Mexico, the city of Teotihuacan ascended at the height of the Classic period; it formed a military and commercial empire whose political influence stretched south into the Maya area and northward. Upon the collapse of Teotihuacán around 600 AD, competition between several important political centers in central Mexico, such as Xochicalco and Cholula, ensued. At this time during the Epi-Classic period, the Nahua peoples began moving south into Mesoamerica from the North, and became politically and culturally dominant in central Mexico, as they displaced speakers of Oto-Manguean languages. During the early post-Classic period, Central Mexico was dominated by the Toltec culture, and Oaxaca by the Mixtec. The lowland Maya area had important centers at Chichén Itzá and Mayapán. Towards the end of the post-Classic period, the Aztecs of Central Mexico built a tributary empire covering most of central Mesoamerica. The distinct Mesoamerican cultural tradition ended with the Spanish conquest in the 16th century. Over the next centuries, Mesoamerican indigenous cultures were gradually subjected to Spanish colonial rule. Aspects of the Mesoamerican cultural heritage still survive among the indigenous peoples who inhabit Mesoamerica. Many continue to speak their ancestral languages, and maintain many practices harking back to their Mesoamerican roots. Etymology and definition The term Mesoamerica literally means "middle America" in Greek. Middle America often refers to a larger area in the Americas, but it has also previously been used more narrowly to refer to Mesoamerica. An example is the title of the 16 volumes of The Handbook of Middle American Indians. "Mesoamerica"