third marriage occurred while he was into his eighties to Anne Twitty, a biologist of Haitian descent. He survived all three of his wives and died in London, May 17, 1991. Beginning of professional career After graduating, he went to Italy to study octopuses. Next he travelled to South Africa where he discovered the field of limnology or the study of freshwater systems, on the shallow lakes near Cape Town. He became an international expert on lakes and wrote a four-volume Treatise on Limnology, with the first volume published in 1957. He took a position teaching zoology at Yale University in 1928. He travelled widely, reaching underexplored parts of the world and writing his first book on the ecology of high-elevation lakes in India. At Yale his graduate students influenced him to research new areas. Research Italy At the age of twenty-two, on graduating from Cambridge, Hutchinson traveled to Italy on a Rockefeller Higher Education Fellowship to work at the Stazione Zoologica. He was interested in doing research on the branchial gland of the octopus. He wanted to establish endocrine function in higher invertebrates. He thought that the branchial gland was the endocrine gland in the octopus, but an octopus shortage put an end to his research. He returned to Italy many times for Italian art, folklore, and to study his Italian ancestry. South Africa In 1926 he applied for a lectureship at the University of Witwatersrand in Johannesburg. He lectured for two years before he was fired, but he continued to study the South African water bugs. When he began his research there were fourteen known species and when he concluded his research there were nearly forty. It was here that Hutchinson discovered limnology, or the study of fresh waters. Along with Grace Pickford, he studied the chemistry and biology of the coastal lakes. He was greatly interested in limnology due to it combining of all his interests such as natural history, aquatic invertebrates, and chemistry. He was drawn to the differences in the chemistry, flora, and fauna in the different water sources. While in South Africa in the late 1920s he frequently visited the southeastern portion of the country, in particular conducting biology and chemistry research in Morgans Bay and St Lucia, he also spent a lot of time around the Sani Pass and the Drakensberg Mountains as well as the area around Royal Natal National Park. He