The Smithsonian Tropical Research Institute (STRI, ) is located in Panama and is the only bureau of the Smithsonian Institution based outside of the United States. It is dedicated to understanding the past, present, and future of tropical ecosystems and their relevance to human welfare. STRI grew out of a small field station established in 1923 on Barro Colorado Island in the Panama Canal Zone to become one of the world's leading tropical research organizations. STRI's facilities provide for long-term ecological studies in the tropics and are used by some 1,200 visiting scientists from academic and research institutions around the world every year. History Smithsonian scientists first came to Panama during the construction of the Panama Canal from 1904 to 1914. The Secretary of the Smithsonian Institution, Charles Doolittle Walcott, reached an agreement with Federico Boyd to conduct a biological inventory of the new Canal Zone in 1910, and this survey was subsequently extended to include all of Panama. Thanks largely to their efforts, the governor of the Canal Zone declared Barro Colorado Island (BCI) a biological reserve in 1923, making it one of the earliest biological reserves in the Americas. During the 1920s and 1930s BCI, in Gatun Lake, became an outdoor laboratory for scientists from U.S. universities and the Smithsonian Institution. By 1940, when BCI was designated the Canal Zone Biological Area (CZBA), more than 300 scientific publications had described the biota of BCI. In the Government Reorganization Act of 1946, BCI became a bureau of the Smithsonian Institution. The Smithsonian Tropical Research Institute (STRI) was created in 1966. With the establishment of STRI, permanent staff scientists were hired and fellowship programs were initiated to support aspiring tropical biologists. The first director after the name change was Martin Humphrey Moynihan. A strong relationship with the Republic of Panama was formalized in the Panama Canal Treaties of 1977, which gradually transferred control of the Canal Zone to Panama. The treaties created the Barro Colorado Nature Monument, which included BCI and five surrounding peninsulas, and granted STRI custody of the monument. Panama granted STRI International Mission status in 1985 and, in 1997, the nation extended STRI's custodianship of facilities beyond the culmination of the Panama Canal Treaties in 1999. STRI's first director after the institute's establishment in 1966 was Martin Humphrey Moynihan. Geographer and scientist Charles F. Bennett, along with his wife, Anna Carole Bennett, participated in STRI during the