and illustrations of monomitic, dimitic and trimitic hyphal construction. He also made major strides in the systematics of clavarioid fungi. As it stood in 1950, the genus Clavaria was a polyphyletic assemblage of coral-shaped fungi. Corner determined that many species of "Clavaria" were actually members of different genera, families and even orders. Using spore characteristics, he determined that a genus of clavarioid fungi—Ramaria—is closely related to a genus of cantharelloid fungi—Gomphus. In his research, Corner stressed the importance of understanding the development and anatomy of fungi, which he realised could not be known just by observing one stage of the specimen. Arguably, his most important contribution on fungal development was determining the role of the basidium as a charged ampoule. He also observed basidiospore development in boletoid and tricholomatoid fungi. In addition to working out systematic relationships of many groups of macrofungi, he also wrote monographs of large groups present in southeast Asia. His work on boletes led to the description of 123 new species and culminated in his classic monograph Boletus in Malaysia as well as a supplement which included species in the genus Phylloporus. Corner also wrote monographs on tricholomatoid, cantharelloid, clavarioid and polyporoid fungi, as well as the genus Thelephora. Corner's monographic work has not only aided those who work in tropical mycology but also those working on the groups in temperate ecosystems. Additionally, these monographs provide the framework for helping mycologists understand the extent of biodiversity in tropical ecosystems. Most of his collections, annotated books and illustrations are housed at the Royal Botanic Garden Edinburgh. His other collections remain at Cambridge or at the United States National Fungus Collection in Beltsville, MD. Personal life In February 1939, Corner married Sheila Kavanagh Bailey in Cambridgeshire. Their first child, John Kavanagh Corner, was born in Singapore in January 1941. The couple also had two daughters, Stephanie Christine Corner born in 1947 in Hanover, New Hampshire, and Dorothy Lindsay Helga Corner in 1948 in Essex. Corner and his wife ultimately divorced in 1952. He remarried in April 1953 to his daughters' nanny, Helga Dinesen Sondergaard (John K Corner). In 1975, he was diagnosed with glaucoma and then cataracts in both eyes, and by 1983 he was unable to do any more microscopy. Even though his physical capacities declined sharply due to muscular paralysis and he was near blindness, Corner's mental faculties remained sharp until his death in 1996. Corner's