with raucous cackling calls, higher pitched and more drawn out than the Manx shearwater's. The yelkouan shearwater has a more contrasted appearance than the Balearic shearwater with which its winter range overlaps, since the latter species is brown above and dirty white below. It is very similar to the black-and-white Manx shearwater of the Atlantic, and stray birds out of their usual range are very difficult to identify with certainty. Also, at least one mixed breeding colony of the yelkouan and the Balearic shearwaters exists on Menorca. A study of these birds recommended that a combination of morphological characteristics and DNA sequence data should be required at least for scientific purposes to assign individual birds to either species. Distribution and habitat Yelkouan shearwaters breed on islands and coastal cliffs in the eastern and central Mediterranean. Most winter in that sea, but small numbers enter the Atlantic in late summer. Behaviour and ecology This species nests in burrows which are only visited at night to avoid predation by large gulls. This is a gregarious species, which can be seen in large numbers from boats or headlands, especially in autumn. The yelkouan shearwater feeds on fish and molluscs. It follows fishing ships when offal is being thrown. Status and conservation It is under some threat from the development of holiday resorts near its breeding sites, and also from animals such as rats and cats. On Le Levant Island, one of its major breeding locations, cats kill thousands of birds each year and it is estimated that this may lead to local extinction in several decades. The study of the Menorcan colony concluded that at least in these westernmost birds, genetic variation was extremely low, suggesting that the yelkouan shearwater may have suffered a marked population decline historically and thus, while not threatened judging from its absolute numbers, it could be vulnerable to adverse effects of inbreeding. It was formerly classified as a species of least concern by the IUCN. But new research has shown it to be rarer than it was believed. Consequently, it was uplisted to near threatened status in 2008 and vulnerable in 2012. References Sources Alcover, Josep Antoni (2001): Nous avenços en el coneixement dels ocells fòssils de les Balears. Anuari Ornitològic de les Balears 16: 3-13. [Article in Catalan, English abstract] PDF fulltext BirdLife International (BLI) (2008): [2008 IUCN Redlist status changes]. Retrieved 2008-MAY-23. Sibley, Charles Gald &