The Crabronidae are a large paraphyletic group (nominally a family) of wasps, including nearly all of the species formerly comprising the now-defunct superfamily Sphecoidea. It collectively includes well over 200 genera, containing well over 9000 species. Crabronids were originally a part of Sphecidae, but the latter name is now restricted to a separate family based on what was once the subfamily Sphecinae. Several of the subfamilies of Crabronidae are often treated as families in their own right, as is true of the most recent phylogenies (example below). Phylogeny This phylogenetic tree is based on Sann et al., 2018, which used phylogenomics to demonstrate that both the bees (Anthophila) and the Sphecidae arose from within the former Crabronidae, which is therefore paraphyletic, and which they suggested should be split into several families; the former family Heterogynaidae nests within the Bembicidae, as here defined. These findings differ in several details from studies published by two other sets of authors in 2017, though all three studies demonstrate a paraphyletic "Crabronidae." Only three of these lineages were not included within Crabronidae in the past: Ampulicidae, Sphecidae, and Anthophila. Subgroups Subfamily Astatinae (now ranked as a family) Astata, etc. Subfamily Bembicinae (now ranked as a family) Tribe Alyssontini Tribe Bembicini Bembix Fabricius 1775 Zyzzyx Pate 1937, etc. Tribe Gorytini Gorytes Latreille, 1804 Sphecius, etc. Tribe Heliocausini Tribe Nyssonini Tribe Stizini Subfamily Crabroninae (incl. Eremiaspheciinae and Dinetinae) Aha Crabro Dalara Liris Pison, etc. Subfamily Mellininae (now ranked as a family) Mellinus Xenosphex Subfamily Pemphredoninae (recently split into three families) Microstigmus Lithium Pemphredon, etc. Subfamily Philanthinae (now ranked as a family) Cerceris Philanthus, etc. Life cycle See also List of Crabronidae genera References Sources Catalog of Sphecidae sensu lato at Cal Academy External links Image Gallery from Gembloux Larra spp., mole cricket hunters on the UF / IFAS Featured Creatures Web site Apocrita families