Liu Oreopteris Holub Tribe Leptogrammeae Todaro 1866 Cyclogramma Tagawa Leptogramma Smith Stegnogramma Blume Tribe Meniscieae Fée 1850-1852 ex Pfeiffer 1873 Subtribe Goniopteridinae Ching 1963 Goniopteris C.Presl Subtribe Menisciinae Payer 1850 Ampelopteris Kunze Cyclosorus Link Meniscium Schreb. Mesophlebion Holttum Subtribe Pseudocyclosorinae Ching 1963 Abacopteris Fée Amblovenatum J.P.Roux Chingia Holttum Christella H.Lév. Glaphyropteridopsis Ching Grypothrix (Holttum 1971) Fawcett & Smith Menisciopsis (Holttum 1982) Fawcett & Smith Menisorus Alston Mesopteris Ching Pakau Fawcett & Smith Parathelypteris (H.Ito) Ching Pelazoneuron (Holttum 1974) Smith & Fawcett Plesioneuron (Holttum) Holttum Pneumatopteris Nakai Pronephrium C.Presl [Nannothelypteris Holttum] Pseudocyclosorus Ching Reholttumia Fawcett & Smith Sphaerostephanos J.Sm. Strophocaulon Fawcett & Smith Trigonospora Holttum Some species of Pronephrium were split off into other genera in 2021 as a result of a phylogenetic study of the family Thelypteridaceae: Abacopteris Fée Grypothrix (Holttum) S.E.Fawc. & A.R.Sm. Menisciopsis (Holttum) S.E.Fawc. & A.R.Sm The extinct genus Holttumopteris from the Cenomanian aged Burmese amber has been suggested to have affinities with the family, but several important diagnostic characters are not preserved. References Fern families