Extinction vortices are a class of models through which conservation biologists, geneticists and ecologists can understand the dynamics of and categorize extinctions in the context of their causes. This model shows the events that ultimately lead small populations to become increasingly vulnerable as they spiral toward extinction. Developed by M. E. Gilpin and M. E. Soulé in 1986, there are currently four classes of extinction vortices. The first two (R and D) deal with environmental factors that have an effect on the ecosystem or community level, such as disturbance, pollution, habitat loss etc. Whereas the second two (F and A) deal with genetic factors such as inbreeding depression and outbreeding depression, genetic drift etc. Types of vortices R Vortex: The R vortex is initiated when there is a disturbance which facilitates a lowering of population size (N) and a corresponding increase in variability (Var(r)). This event can make populations vulnerable to additional disturbances which will lead to further decreases in population size (N) and further increases in variability (Var(r)). A prime example of this would be the disruption of sex ratios in a population away from the species optimum. D Vortex: The D vortex is initiated when population size (N) decreases and variability (Var(r)) increases such that the spatial distribution (D) of the population is increased and the population becomes "patchy" or fragmented. Within these fragments, local extinction rates increase which, through positive feedback, further increases D. F Vortex: The F vortex is initiated by a decrease in population size (N) which leads to a decrease in heterozygosity, and therefore a decrease in genetic diversity. Decreased population size makes the effects of genetic drift more prominent, resulting in increased risk of inbreeding depression and an increase in population genetic load, which over time will result in extinction. A Vortex: The A vortex is a result of an increase in the impact of genetic drift on the population, due to the population's decreased size. This corresponds with a decrease in genetic variance which leads to a decrease in "population adaptive potential", and eventual extinction. This vortex can result from biological invasion, resulting in large scale hybridization and outbreeding depression. Extinction vortex factors Environmental factors Many of the environmental events that contribute to an extinction vortex do so through reduction in population size. These events can include rapid loss of population size due to disease, natural disasters, and climate change. Habitat loss