skin. Etymology The mantled guereza has many alternative common names including the guereza, the eastern black-and-white colobus, the magistrate colobus, or the Abyssinian black-and-white colobus. The name "mantled" refers to its mantle, the long silky white fringes of hair that run along its body and "guereza" is the native name of the monkey in Ethiopia. The scientific name Colobus derives from Greek kolobus meaning "mutilated" which refers to its lack of thumbs. Taxonomic classification The mantled guereza was first classified by Eduard Rüppell, a German naturalist and explorer, during his trip to Abyssinia between 1830 and 1834. He wrote about the species in Neue Wirbelthiere con Abyssinien, Saengthiere in 1835. It was first seen in Europe in 1890 in Berlin Zoological Garden when three individuals were purchased from a dealer from Massawa, Eritrea. The mantled guereza is in the Colobinae subfamily, also known as the leaf-eating monkeys, a group of Old World monkeys from Asia and Africa. This subfamily is split into three groups, the colobus monkeys of Africa, of which the mantled guereza is a part, the langurs, or leaf monkeys, of Asia, and an "odd-nosed" group. The African colobus monkeys are divided again by distinctions in color, behavior, and ecology. The three genera are the black-and-white colobi, the red colobi, and the olive colobi. There are three black-and-white colobi: the mantled guereza, Colobus guereza, the king colobus, C. polykomos, and the Angola colobus, C. angolensis. Groves lists seven subspecies of mantled guereza in Mammal Species of the World (MSW) (2005). The validity of the Djaffa Mountain guereza, C. g. gallarum, is uncertain, although not listed by Groves in MSW, it is recognised in his 2007 Colobinae review paper, and by International Union for Conservation of Nature (IUCN) assessors Gippolliti and Butynski in 2008. Western guereza, Colobus guereza occidentalis, occurs from eastern Nigeria, Cameroon, and Gabon at the edge of its western range to South Sudan and Uganda, west of the Nile. Omo River guereza or Abyssinian black-and-white colobus, C. g. guereza, found in Ethiopia, in the highlands west of the Rift Valley down to the reaches of the Awash River, the Omo River, and in the Blue Nile gorge. Djaffa Mountains guereza or Neumann's black-and-white colobus, C. g. gallarum, found in the Ethiopian Highlands east of the Rift Valley. Dodinga Hills guereza, C. g. dodingae, found in the Didinga Hills in South Sudan. Mau Forest guereza, C. g.