1770 but any specimens collected were lost later in the voyage. The first formal description of a greenhood was by Jacques Labillardière who collected a specimen from Bruny Island and gave it the name Disperis alata. After several name changes, Disperis alata is now known as Pterostylis alata. John White, the first surgeon-general of the colony of New South Wales collected several species and George Caley collected 208 greenhood specimens from 16 species and described them in great detail, including details of their habitat and gave them names like Druids Cap patersoni but did not publish his work. The genus Pterostylis was first raised by Robert Brown who formally described 19 species but did not nominate a type species. The species he described were collected from the east coast of the mainland and from Tasmania and the descriptions were published in Prodromus Florae Novae Hollandiae. Alan Cunningham was the first to describe a New Zealand species, P. banksii and John Lindley described the first four Western Australian species, P. vittata, P. pyramidalis, P. barbata and P. scabra from specimens collected by James Drummond. About 300 species are recognised and about 200 of them have been formally described and named. In 2001 the genus Pterostylis was split such that two new genera, Plumatichilos and Oligochaetochilus, were established based on morphological characters. These divisions effectively separated species with a thread-like, feathery labellum (Plumatichilos), and those with downward-curved lateral sepals (Oligochaetochilus) from the remaining Pterostylis species. The following year, David Jones and Mark Clements proposed further separating Pterostylis into sixteen genera based on a combination of molecular and morphological characters: the genera Bunochilus, Crangonorchis, Diplodium, Eremorchis, Hymenochilus, Linguella, Oligochaetochilus, Petrorchis, Pharochilum, Plumatichilos, Ranorchis, Speculantha, Stamnorchis, Taurantha and Urochilus. These changes were not adopted widely by Australian herbaria as the data appeared to support a monophyletic group. In 2010, further molecular work reinstated Pterostylis as a monophyletic group with two subgenera distinguishing species with lateral sepals that are bent sharply downwards ("deflexed") from those with the lateral sepals bent backwards ("recurved"). The genus name (Pterostylis) is derived from the Greek words pteron meaning 'wing' and stylos meaning 'pillar' or 'post', but in orchids, generally applies to the column. The closest relative of Pterostylis is Achlydosa, restricted to New Caledonia and the sole other genus of subtribe Pterostylidinae. Distribution and habitat Orchids in the genus Pterostylis occur mostly in Australia but are also found in