was lower than that of Palaeopropithecus. Behavior Archaeoindris is thought to have been a leaf-eater (folivorous), a view supported by wear patterns on its teeth. Its fused mandibular symphyses and the likelihood of accelerated dental development suggest that it began processing fibrous foods in the mouth with thorough chewing. Its diet may also have included some fruits and seeds. Like most of the other giant lemurs, Archaeoindris is thought to have been diurnal because of its relatively small orbits, which are comparable to those of gorillas. Both Standing and Lamberton assumed Archaeoindris to be a slow-moving tree-dweller like Megaladapis, primarily due to the misattributed bones. Lamberton also speculated that it would have resembled a ground sloth—a view later supported by Jungers in 1980 after several misattributions had been corrected and having considered its gorilla-like size. Jungers went on to propose that it would have spent most of its time on the ground (terrestrial). However, the functional morphology of its hip joint indicates a degree of mobility typically seen in more arboreal animals. Other traits shared with Palaeopropithecus, particularly seen in the femur, suggest that Archaeoindris spent considerable time in the trees for feeding and possibly nesting, although it also would have visited the ground to feed and travel. It is described as a deliberate, scansorial (climbing) browser, and it is unknown whether it was like Palaeopropithecus in performing hang-feeding since hand and foot bones are missing. Given its bulky size, this would be unexpected. Distribution and habitat Archaeoindris is known from only one subfossil site, Ampasambazimba, in central Madagascar, and all remains date to the Late Quaternary. The area today is dominated by grasslands, particularly of the grass genus Aristida. Prior to human arrival, the area around Ampasambazimba was not completely forested, but more of an open habitat, consisting of a mix of woodlands, bushlands, and savanna. Animal remains at this subfossil site have yielded about 20 species of lemur living in sympatry (sharing the same geographic area). In comparison, the nearby Ambohitantely Special Reserve today contains only four species, roughly 20% of the area's original lemur diversity. Extinction Despite being the most species-rich family among the giant lemurs, all four genera of sloth lemur, including Archaeoindris, have gone extinct. Radiocarbon dating of the stratigraphic level of some of the Archaeoindris remains were dated to 8000 BP, while two other specimens were dated to 2362–2149 BP (412–199 BCE) and 2711–2338