in a flora of Brazil, Engler described many new species in Ochnaceae, especially in its largest genus, Ouratea. He described 85 species in Ouratea, 17 of which he named as new species at that time. He also transferred 63 species to Ouratea from other genera. The genera Quiina and Touroulia had been known since 1775, when they were described by Jean Baptiste Aublet, and they were variously classified by 19th century taxonomists. Jacques Denys Choisy erected the family Quiinaceae (as Quiinacées) for them in 1849, but he did not meet the requirements for valid publication of a botanical name. The name Quiinaceae was validated by Engler in Flora Brasiliensis in 1888. The genus Medusagyne had been described by John Gilbert Baker in 1877, in a flora of Mauritius and the Seychelles, but it was not until 1924 that it was segregated into its own monogeneric family. In 1893, Ernest Friedrich Gilg covered Ochnaceae, and Adolf Engler covered Quiinaceae for the first edition of Die Natürlichen Pflanzenfamilien. Engler wrote a description of Medusagyne in a supplement to the first edition of DNP in 1897. He placed Medusagyne under the heading "Zweifelhafte, möglicherweise zu den Guttiferae gehörige Gattung" (Doubtful, possibly a genus belonging to Guttiferae). Guttiferae is an obsolete name for Clusiaceae. In 1902, Philippe van Tieghem recognized six families in what is now the subfamily Ochnoideae. These were Luxemburgiaceae, Sauvagesiaceae, Wallaceaceae, Euthemidaceae, Lophiraceae, and Ochnaceae. Three of these, (Wallaceaceae, Euthemidaceae, and Lophiraceae) were monogeneric and were erected by van Tieghem at that time. His Luxemburgiaceae included the basal clades of what is now the tribe Sauvagesieae. Van Tieghem named a great many genera in 1902, circumscribing them very narrowly. In what is now the subtribe Ochninae, he delineated 53 genera. The most recent revision of that group divides it into six genera. In 1925, for the second edition of DNP, Engler and Gilg expanded on their treatment of Quiinaceae and Ochnaceae, respectively, compared to what they had written in 1893. Medusagynaceae was covered in the same volume of DNP by Adolf Engler and Hans Melchior. For the second edition of DNP, Engler recognized two genera, Quiina and Touroulia, in Quiinaceae. Lacunaria and Froesia were discovered later and named in 1925 and 1948, respectively. In that same volume, Gilg divided his Ochnaceae (equivalent to modern Ochnoideae) into 21 genera, including Indovethia, Leitgebia, Vausagesia, and Lauradia (as Lavradia), which are now regarded as