and the other proposes only presynaptic changes, graded ability, and age-dependence. These theories do agree on one element of LTP, namely, that it must occur through physical changes to the synaptic membrane/s, i.e. synaptic plasticity. Perceptual control theory encompasses both of these views. It proposes the mechanism of 'reorganisation' as the basis of learning. Reorganisation occurs within the inherent control system of a human or animal by restructuring the inter- and intraconnections of its hierarchical organisation, akin to the neuroscientific phenomenon of neural plasticity. This reorganisation initially allows the trial-and-error form of learning, which is seen in babies, and then progresses to more structured learning through association, apparent in infants, and finally to systematic learning, covering the adult ability to learn from both internally and externally generated stimuli and events. In this way, PCT provides a valid model for learning that combines the biological mechanisms of LTP with an explanation of the progression and change of mechanisms associated with developmental ability. Powers in 2008 produced a simulation of arm co-ordination. He suggested that in order to move your arm, fourteen control systems that control fourteen joint angles are involved, and they reorganise simultaneously and independently. It was found that for optimum performance, the output functions must be organised in a way so as each control system's output only affects the one environmental variable it is perceiving. In this simulation, the reorganising process is working as it should, and just as Powers suggests that it works in humans, reducing outputs that cause error and increasing those that reduce error. Initially, the disturbances have large effects on the angles of the joints, but over time the joint angles match the reference signals more closely due to the system being reorganised. Powers suggests that in order to achieve coordination of joint angles to produce desired movements, instead of calculating how multiple joint angles must change to produce this movement the brain uses negative feedback systems to generate the joint angles that are required. A single reference signal that is varied in a higher-order system can generate a movement that requires several joint angles to change at the same time. Hierarchical organisation Botvinick in 2008 proposed that one of the founding insights of the cognitive revolution was the recognition of hierarchical structure in human behavior. Despite decades of research, however, the computational mechanisms underlying hierarchically organized behavior are still not fully understood. Bedre, Hoffman,