Cooney & D'Esposito in 2009 proposed that the fundamental goal in cognitive neuroscience is to characterize the functional organization of the frontal cortex that supports the control of action. Recent neuroimaging data has supported the hypothesis that the frontal lobes are organized hierarchically, such that control is supported in progressively caudal regions as control moves to more concrete specification of action. However, it is still not clear whether lower-order control processors are differentially affected by impairments in higher-order control when between-level interactions are required to complete a task, or whether there are feedback influences of lower-level on higher-level control. Botvinik in 2008 found that all existing models of hierarchically structured behavior share at least one general assumption – that the hierarchical, part–whole organization of human action is mirrored in the internal or neural representations underlying it. Specifically, the assumption is that there exist representations not only of low-level motor behaviors, but also separable representations of higher-level behavioral units. The latest crop of models provides new insights, but also poses new or refined questions for empirical research, including how abstract action representations emerge through learning, how they interact with different modes of action control, and how they sort out within the prefrontal cortex (PFC). Perceptual control theory (PCT) can provide an explanatory model of neural organisation that deals with the current issues. PCT describes the hierarchical character of behavior as being determined by control of hierarchically organized perception. Control systems in the body and in the internal environment of billions of interconnected neurons within the brain are responsible for keeping perceptual signals within survivable limits in the unpredictably variable environment from which those perceptions are derived. PCT does not propose that there is an internal model within which the brain simulates behavior before issuing commands to execute that behavior. Instead, one of its characteristic features is the principled lack of cerebral organisation of behavior. Rather, behavior is the organism's variable means to reduce the discrepancy between perceptions and reference values which are based on various external and internal inputs. Behavior must constantly adapt and change for an organism to maintain its perceptual goals. In this way, PCT can provide an explanation of abstract learning through spontaneous reorganisation of the hierarchy. PCT proposes that conflict occurs between disparate reference values for a given perception rather than between different responses, and that learning is implemented as trial-and-error changes of the properties of control