who was Sutton's teacher and Boveri's friend, called this the "Sutton-Boveri Theory". Between 1902 and 1904 Theodor Heinrich Boveri (1862–1915), a German biologist, made several contributions to chromosome theory in a series of papers, finally stating in 1904 that he had seen the link between chromosomes and Mendel's results in 1902 (although this is not documented in his publications). He said that chromosomes were "independent entities which retain their independence even in the resting nucleus... What comes out of the nucleus is what goes into it". Works An Introduction to General Biology (1887), with W. T. Sedgwick The Embryology of the Earthworm (1889) Amphioxus, and the Mosaic Theory of Development (1893) Atlas of Fertilization and Karyokinesis (1895) The Cell in Development and Inheritance (1896; second edition, 1915; third edition, 1925) The Physical Basis of Life (1923) References Bibliography Al-Awqati, Q. 2002. Edmund Beecher Wilson: America's First Cell Biologist. Living Legacies, Columbia University. Gilbert, S. F. 2003. Edmund Beecher Wilson and Frank R. Lillie and the relationship between evolution and development, Developmental Biology, Seventh edition, Sinauer External links 1856 births 1939 deaths American science writers American geneticists American zoologists Johns Hopkins University alumni Yale University alumni Members of the United States National Academy of Sciences Foreign Members of the Royal Society Members of the Royal Netherlands Academy of Arts and Sciences Fellows of the American Academy of Arts and Sciences People from Geneva, Illinois Members of the American Philosophical Society