83 million years ago in the Cretaceous within an entomophilic clade within Moraceae that includes tribe Castilleae and genus Ficus, as the bracts protecting the inflorescence tightened to form the ostiole. This greatly increased the pollinator specificity of the plant and initiated a long and complex history of coevolution between figs and their pollinating wasps (agaonids). References Fruit morphology