2019 was reinterpreted as an early diverging relative of the family Hydroscaphidae. A fossil impression assigned to the living genus Hydroscapha in Hydroscaphidae is known from the Yixian Formation in the Jehol Biota, dating from the Early Cretaceous (Aptian). Fossils from the early Late Cretaceous (Cenomanian) aged Burmese amber from Myanmar, have been assigned to the extant genus Lepicerus and extinct genus Lepiceratus within Lepiceridae, as well as the extant Sphaerius and Bezesporum and the extinct Burmasporum and Crowsonaerius, belonging to Sphaerusidae. Gallery See also List of subgroups of the order Coleoptera Notes References External links Insect suborders Beetles described in 1955 Beetles of North America Beetles of Asia