followed the Dvina, Vologda and Rybinsk basins respectively. Lobes originated as result of ice following shallow topographic depressions filled with a soft sediment substrate. Permafrost covered Europe south of the ice sheet down to as far south as present-day Szeged in Southern Hungary. Ice covered the whole of Iceland. In addition, ice covered Ireland and almost all of Wales, with the southern boundary of the ice sheet running approximately from the current location of Cardiff north-north-east to Middlesbrough, and then across the now submerged land of Doggerland to Denmark. In the Cantabrian Mountains of the northwestern corner of the Iberian Peninsula, which in the present day have no permanent glaciers, the LGM led to a local glacial recession as a result of increased aridity caused by the growth of other ice sheets farther to the east and north, which drastically limited annual snowfall over the mountains of northwestern Spain. The Cantabrian alpine glaciers had previously expanded between approximately 60,000 and 40,000 years ago during a local glacial maximum in the region. North America In North America, the Laurentide Ice Sheet grew rapidly at the onset of the LGM until it covered essentially all of Canada east of the Rocky Mountains and extended roughly to the Missouri and Ohio Rivers, and eastward to Manhattan, reaching a total maximum volume of around 26.5 to 37 million cubic kilometres. In addition to the large Cordilleran Ice Sheet in Canada and Montana, alpine glaciers advanced and (in some locations) ice caps covered much of the Rocky and Sierra Nevada Mountains further south. Latitudinal gradients were so sharp that permafrost did not reach far south of the ice sheets except at high elevations. Glaciers forced the early human populations who had originally migrated from northeast Siberia into refugia, reshaping their genetic variation by mutation and drift. This phenomenon established the older haplogroups found among Native Americans, and later migrations are responsible for northern North American haplogroups. On the Island of Hawaii, geologists have long recognized deposits formed by glaciers on Mauna Kea during recent ice ages. The latest work indicates that deposits of three glacial episodes since 150,000 to 200,000 years ago are preserved on the volcano. Glacial moraines on the volcano formed about 70,000 years ago and from about 40,000 to 13,000 years ago. If glacial deposits were formed on Mauna Loa, they have long since been buried by younger lava flows. South America