The Berothidae are a family of winged insects of the order Neuroptera. They are known commonly as the beaded lacewings. The family was first named by Anton Handlirsch in 1906. The family consists of 24 genera and 110 living species distributed discontinuously worldwide, mostly in tropical and subtropical regions. Numerous extinct species have also been described. Their ecology is poorly known, but in the species where larval stages have been documented, the larvae are predators of termites. Systematics The peculiar genus Lomamyia cannot be robustly assigned to any one subfamily. In addition, a considerable fossil diversity of beaded lacewings is known from the Late Jurassic onwards, containing numerous genera which are likewise basal or incertae sedis. Extant genera Asadeteva Aspöck & Aspöck, 1981 Austroberothella Aspöck & Aspöck, 1985 Berlekrumyia Aspöck & Aspöck, 1988 Berotha Walker, 1860 Berothimerobius Monserrat & Deretsky, 1999 Cyrenoberotha MacLeod & Adams, 1968 Isoscelipteron Costa, 1863 Lekrugeria Navás, 1929 Lomamyia Banks, 1904 Manselliberotha Aspöck & Aspöck, 1988 Naizema Navás, 1919 Nodalla Navás, 1926 Nosybus Navás, 1910 Nyrma Navás, 1933 Ormiscocerus Blanchard, 1851 Podallea Navás, 1936 Protobiella Tillyard, 1923 Quasispermophorella Aspöck & Aspöck, 1986 Spermophorella Tillyard, 1916 Spiroberotha Adams, 1990 Stenobiella Tillyard, 1916 Tanzanberotha Aspöck & Hynd, 1995 Trichoberotha Handschin, 1935 Trichoma Tillyard, 1916 Extinct genera †Ansoberotha Yang et al. 2019 Burmese amber, Myanmar, Late Cretaceous (Cenomanian) †Araripeberotha Martins-Neto and Vulcano 1990 Crato Formation, Brazil, Early Cretaceous (Aptian) †Banoberotha Whalley 1980 Lebanese amber, Early Cretaceous (Barremian) †Caririberotha Martins-Neto and Vulcano 1990 Crato Formation, Brazil, Aptian †Cantabroberotha Pérez-de la Fuente, Peñalver & Engel, 2020, Spanish amber, Early Cretaceous (Albian) †Cornoberotha Yang et al. 2019 Burmese amber, Myanmar, Cenomanian †Dasyberotha Engel and Grimaldi 2008 Burmese amber, Myanmar, Cenomanian †Elektroberotha Makarkin and Ohl 2015 Baltic amber, Eocene †Epimesoberotha Jepson et al. 2012 Lulworth Formation, United Kingdom, Early Cretaceous (Berriasian) †Ethiroberotha Engel and Grimaldi 2008 Burmese amber, Myanmar, Cenomanian †Haploberotha Engel and Grimaldi 2008 Burmese amber, Myanmar, Cenomanian †Iceloberotha Engel and Grimaldi 2008 Burmese amber, Myanmar, Cenomanian †Jersiberotha Grimaldi 2000 Burmese amber, Myanmar, Cenomanian New Jersey amber, Late Cretaceous (Turonian) †Krokhathone Khramov 2015 Karabastau Formation, Kazakhstan, Middle/Late Jurassic (Callovian/Oxfordian) †Maculaberotha Yuan et al. 2016 Burmese amber, Myanmar, Cenomanian †Magniberotha Yuan et al. 2016 Burmese amber, Myanmar, Cenomanian †subfamily Mesithoninae Panfilov 1980 †Berothone Khramov 2015 Karabastau Formation, Kazakhstan, Callovian/Oxfordian †Mesithone Panfilov 1980 Karabastau Formation, Kazakhstan, Callovian/Oxfordian Zaza Formation, Turga Formation, Russia, Aptian †Pseudosisyra Makarkin 1999 Zaza Formation, Russia, Aptian †Microberotha Archibald and Makarkin