wings (Coniopterygidae). And even though they are not often placed in the superfamily Coniopterygoidea yet, they most likely form a clade with the Coniopterygidae and thus it would seem that the Coniopterygoidea, rather than being an unnecessarily monotypic taxon, is expanded to signify that the spongillaflies and the dusty wings are each other's closest relatives. A few fossil genera are known, mainly from the Eocene like "Sisyra" amissa which may or may not be the first record of the living genus. The genus Cratosisyrops, from the Aptian Crato Formation, was suggested to be a spongillafly member, but later authors Perkovsky and Makarkin in 2015 noted the fossil lacks details to confidently place it in the family. The oldest members of the family are species in the subfamily Paradoxosisyrinae from Cenomanian Burmese amber and Prosisyrina from Santonian Taimyr amber. The family is currently divided into two subfamilies with the four extant genera and two of the extinct genera placed into Sisyrinae, while five extinct genera are placed into the subfamily Paradoxosisyrinae. Sisyridae †Stictosisyra Yang et al. 2018 Burmese amber, Myanmar, Late Cretaceous (Cenomanian) †Paradoxosisyrinae Burmese amber, Myanmar, Late Cretaceous (Cenomanian) †Buratina †Khobotun †Paradoxosisyra †Protosiphoniella †Sidorchukatia Sisyrinae Climacia †Paleosisyra Oise amber, France, Eocene (Ypresian) Baltic amber, Eocene Bembridge Marls, United Kingdom, Eocene (Priabonian) †Prosisyrina Taimyr amber, Russia, Late Cretaceous (Santonian) Sisyborina Sisyra Sisyrina References This article draws heavily on the corresponding article in the German-language Wikipedia. Neuroptera Coniopterygoidea Neuroptera families