published their Genera plantarum (1862–1883) ordo Amaryllideae contained five tribes, and tribe Amarylleae 3 subtribes (see Bentham & Hooker system). They placed Galanthus in subtribe Genuinae and included three species. Phylogeny Galanthus is one of three closely related genera making up the tribe Galantheae within subfamily Amaryllidoideae (family Amaryllidaceae). Sometimes snowdrops are confused with the other two genera, Leucojum and Acis (both called snowflakes). Leucojum species are much larger and flower in spring (or early summer, depending on the species), with all six tepals in the flower being the same size, although some "poculiform" (goblet- or cup-shaped) Galanthus species may have inner segments similar in shape and length to the outer ones. Galantheae are likely to have arisen in the Caucusus. Subdivision Galanthus has approximately 20 species, but new species continue to be described. G. trojanus was identified in Turkey in 2001. G. panjutinii (Panjutin's snowdrop) was discovered in 2012 in five locations in a small area (estimated at ) of the northern Colchis area (western Transcaucasus) of Georgia and Russia. G. samothracicus was identified in Greece in 2014. Since it has not been subjected to genetic sequencing, it remains unplaced. It resembles G. nivalis, but is outside the distribution of that species. Many species are difficult to identify, however, and traditional infrageneric classification based on plant morphology alone, such as those of Stern (1956), Traub (1963) and Davis (1999, 2001), has not reflected what is known about its evolutionary history, due to the morphological similarities among the species and relative lack of easily discernible distinguishing characteristics. Stern divided the genus into three series according to leaf vernation (the way the leaves are folded in the bud, when viewed in transverse section, see Description); section Nivales Beck (flat leaves) section Plicati Beck (plicate leaves) section Latifolii Stern (convolute leaves) Stern further utilised characteristics such as the markings of the inner segments, length of the pedicels in relation to the spathe, and the colour and shape of the leaves in identifying and classifying species Traub considered them as subgenera; subgenus Galanthus subgenus Plicatanthus Traub & Moldk. subgenus Platyphyllanthe Traub By contrast Davis, with much more information and specimens, included biogeography in addition to vernation, forming two series. He used somewhat different terminology for vernation, namely applanate (flat), explicative (plicate), and supervolute (convolute). He merged Nivalis and Plicati into series Galanthus, and divided Latifolii into two subseries, Glaucaefolii (Kem.-Nath) A.P.Davis and Viridifolii