and Dugandiodendron, which are then placed in subgenus Magnolia, and genus Manglietia, which could be joined with subgenus Magnolia or may even earn the status of an extra subgenus. Elmerrillia seems to be closely related to Michelia and Yulania, in which case it will most likely be treated in the same way as Michelia is now. The precise nomenclatural status of small or monospecific genera like Kmeria, Parakmeria, Pachylarnax, Manglietiastrum, Aromadendron, Woonyoungia, Alcimandra, Paramichelia and Tsoongiodendron remains uncertain. Taxonomists who merge Michelia into Magnolia tend to merge these small genera into Magnolia s.l. as well. Botanists do not yet agree on whether to recognize a big Magnolia or the different small genera. For example, Flora of China offers two choices: a large genus Magnolia which includes about 300 species, everything in the Magnoliaceae except Liriodendron (tulip tree), or 16 different genera, some of them recently split out or re-recognized, each of which contains up to 50 species. The western co-author favors the big genus Magnolia, whereas the Chinese recognize the different small genera. Fossil record Fossils assignable to Magnolia extend into the Paleogene, such as Magnolia nanningensis, named for mummified wood from the Oligocene of Guangxi, China, which has a close affinity to members of the modern secton Michelia. Subdivision In 2012, the Magnolia Society published on its website a classification of the genus produced by Richard B. Figlar, based on a 2004 classification by Figlar and Hans Peter Nooteboom. Species of Magnolia were listed under three subgenera, 12 sections, and 13 subsections. Subsequent molecular phylogenetic studies have led to some revisions of this system; for example, the subgenus Magnolia was found not to be monophyletic. A revised classification in 2020, based on a phylogenetic analysis of complete chloroplast genomes, abandoned subgenera and subsections, dividing Magnolia into 15 sections. The relationships among these sections are shown in the following cladogram, as is the paraphyletic status of subgenus Magnolia. The table below compares the 2012 and 2020 classifications. (The exact circumscriptions of the corresponding taxa may not be the same.) Species The species lists below are divided according to the Magnolia Society's 2012 classification. Subgenus Magnolia Anthers open by splitting at the front facing the centre of the flower, deciduous or evergreen, flowers produced after the leaves. Section Magnolia Magnolia faustinomirandae A.Vázquez - (Mexico) Magnolia grandiflora L. - (SE US) Magnolia guatemalensis Donn. Sm. - (Guatemala, Honduras, El Salvador) Magnolia guatemalensis