a lecture, naming the new genus Dryandra in 1809, however Joseph Knight published the name Josephia before Brown published his paper with the description of Dryandra. Brown ignored Knight's name, as did subsequent botanists. In 1891, Otto Kuntze, strictly applying the principle of priority, argued that Pimelea should revert to the name Banksia J.R.Forst & G.Forst. He proposed the new genus Sirmuellera to replaced Banksia L.f. and transferred its species to the new genus. This arrangement was largely ignored by Kuntze's contemporaries.Banksia L.f. was formally conserved and Sirmuellera rejected in 1940. Banksia belongs to the family Proteaceae, subfamily Grevilleoideae, and tribe Banksieae. There are around 170 species. The closest relatives of Banksia are two genera of rainforest trees in North Queensland (Musgravea and Austromuellera). Subgeneric arrangement Alex George arranged the genus into two subgenera—subgenus Isostylis (containing B. ilicifolia, B. oligantha and B. cuneata) and subgenus Banksia (containing all other species except those he considered dryandras)—in his 1981 monograph and 1999 treatment for the Flora of Australia series. He held that flower morphology was the key to relationships in the genus. Austin Mast and Kevin Thiele published the official merging of Dryandra within Banksia in 2007, recalibrating the genus into subgenus Banksia and subgenus Spathulatae. Distribution and habitat All but one of the living Banksia species are endemic to Australia. The exception is B. dentata (tropical banksia), which occurs throughout northern Australia, and on islands to the north including New Guinea and the Aru Islands. An extinct species, B. novae-zelandiae, was found in New Zealand. The other species occur in two distinct geographical regions: southwest Western Australia and eastern Australia. Southwest Western Australia is the main centre of biodiversity; over 90% of all Banksia species occur only there, from Exmouth in the north, south and east to beyond Esperance on the south coast. Eastern Australia has far fewer species, but these include some of best known and most widely distributed species, including B. integrifolia (coast banksia) and B. spinulosa (hairpin banksia). Here they occur from the Eyre Peninsula in South Australia right around the east coast up to Cape York in Queensland. The vast majority of Banksia are found in sandy or gravelly soils, though some populations of B. marginata (silver banksia) and B. spinulosa do occur on soil that is heavier and more clay-like. B. seminuda is exceptional for its preference for rich loams along watercourses. Most occur in heathlands