The enhanced nitrification-denitrification coupling contributes to greater removal of biologically available nitrogen in shallow and coastal environments, which can be further enhanced by the excretion of ammonium by bioturbators and other organisms residing in bioturbator burrows. While both nitrification and denitrification are enhanced by bioturbation, the effects of bioturbators on denitrification rates have been found to be greater than that on rates of nitrification, further promoting the removal of biologically available nitrogen. This increased removal of biologically available nitrogen has been suggested to be linked to increased rates of nitrogen fixation in microenvironments within burrows, as indicated by evidence of nitrogen fixation by sulfate-reducing bacteria via the presence of nifH (nitrogenase) genes. Bioturbation by walrus feeding is a significant source of sediment and biological community structure and nutrient flux in the Bering Sea. Walruses feed by digging their muzzles into the sediment and extracting clams through powerful suction. By digging through the sediment, walruses rapidly release large amounts of organic material and nutrients, especially ammonium, from the sediment to the water column. Additionally, walrus feeding behavior mixes and oxygenates the sediment and creates pits in the sediment which serve as new habitat structures for invertebrate larvae. Deep-sea Bioturbation is important in the deep sea because deep-sea ecosystem functioning depends on the use and recycling of nutrients and organic inputs from the photic zone. In low energy regions (areas with relatively still water), bioturbation is the only force creating heterogeneity in solute concentration and mineral distribution in the sediment. It has been suggested that higher benthic diversity in the deep sea could lead to more bioturbation which, in turn, would increase the transport of organic matter and nutrients to benthic sediments. Through the consumption of surface-derived organic matter, animals living on the sediment surface facilitate the incorporation of particulate organic carbon (POC) into the sediment where it is consumed by sediment dwelling animals and bacteria. Incorporation of POC into the food webs of sediment dwelling animals promotes carbon sequestration by removing carbon from the water column and burying it in the sediment. In some deep-sea sediments, intense bioturbation enhances manganese and nitrogen cycling. Role in Organic Contaminant Flux Bioturbation can either enhance or reduce the flux of contaminants from the sediment to the water column, depending on the mechanism of sediment transport. In polluted sediments, bioturbating animals can mix the surface layer and cause the release of sequestered contaminants into the