UK population geneticist, lepidopterist, human blood group researcher G. H. Shull (1874–1954), US geneticist, made key discoveries including heterosis Torsten Sjögren Mark Skolnick (born 1946), US geneticist, developed Restriction Fragment Length Polymorphisms (RFLPs) for genetic mapping, and founded Myriad Genetics Obaid Siddiqi (1932–2013), Indian neurogeneticist, pioneer on olfactory sense of fruit fly Drosophila David Sillence (born 1944), Australian clinical geneticist, pioneered training of Australian geneticists, research in bone dysplasias, classified osteogenesis imperfecta Norman Simmons (1915–2004), US, forgotten donor of pure DNA to Rosalind Franklin in double helix saga Piotr Słonimski (1922–2009), Polish-Parisian yeast geneticist, pioneer of mitochondrial heredity William S. Sly (born 1932), US biochemical geneticist, mucopolysaccharidosis type VII (Sly syndrome) Cedric A. B. Smith (1917–2002), British statistician, made key contributions to statistical genetics David W. Smith (1926–1981), US pediatrician, influential dysmorphologist, named fetal alcohol syndrome Hamilton Smith (born 1931), US microbiologist, Nobel Prize for restriction endonucleases Michael Smith (1932–2000), UK-born Canadian biochemist, Nobel Prize for site-directed mutagenesis Oliver Smithies (1925–2017), UK/US molecular geneticist, inventor, gel electrophoresis, knockout mice George Snell (1903–1996), US mouse geneticist, pioneer transplant immunologist, won Nobel Prize Lawrence H. Snyder (1901–1986), US pioneer in medical genetics, studied blood groups Robert R. Sokal (1925–2012), Austrian-born US biological anthropologist and biostatistician. Tracy M. Sonneborn (1905–1981), protozoan biologist and geneticist Ed Southern (born 1938), UK molecular biologist, invented Southern blot and DNA microarray technologies Hans Spemann (1869–1941) German embryologist, Nobel Prize for discovery of embryonic induction David Stadler, US geneticist, mechanisms of mutation and recombination in Neurospora L. J. Stadler (1896–1954), eminent Us maize geneticist, father of David Stadler Frank Stahl (born 1929), US molecular biologist, the Stahl half of the Meselson-Stahl experiment David States, US geneticist and bioinformatician, computational study of human genome and proteome G. Ledyard Stebbins (1906–2000), US botanist, geneticist and evolutionary biologist Michael Stebbins, US geneticist, science writer, editor and activist Emmy Stein (1879–1954), German botanist and geneticist Joan A. Steitz (born 1941), US molecular biologist, pioneering studies of snRNAs and snRNPs (snurps) Gunther Stent (1924–2008), German-born US molecular geneticist, phage worker, philosopher of science Curt Stern (1902–1981), German-born US Drosophila and human geneticist, great teacher Nettie Stevens (1861–1912), US geneticist, studied chromosomal basis of sex and discovered XY basis Miodrag Stojković (born 1964), Serbian geneticist, working in Europe on mammalian cloning George Streisinger (1927–1984), US geneticist, work on bacterial viruses, frameshift mutations Leonell Strong (1894–1982), US geneticist, mouse geneticist and cancer researcher Alfred Sturtevant