ended with Andrey being thrown into a cell full of hungry dogs and devoured by them (1543). In 1540, Metropolitan Joasaphus managed to recall Ivan Belsky from exile, helping him clear the court of the Shuyskys. Two years later, Ivan Shuysky instigated a military revolt and again gained power. He had Macarius elected the new metropolitan and regent, but Macarius gradually ousted him from the Kremlin and persuaded him to resign his powers. Ivan Vasilevich Shuysky died in semi-obscurity in 1546. Military heroes Andrey Mikhailovich's elder brother, Prince Ivan Mikhailovich Pleten', was one of the leading Muscovite generals between 1531 and his death in 1559. During the regency of Elena Glinskaya he served as the governor of Moscow and of Kholmogory. In 1540, he was put in charge of the Russian army operating in Livonia. In 1542 he routed the Crimean Tatars. Two years later, he was recorded as operating against Kazan. In the late 1540s, he administrated the royal palaces. In 1553, Ivan Pleten' signed an armistice with the Grand Duchy of Lithuania. During the later part of Ivan IV's reign, the Shuyskys stood aloof from the macabre politics of the Oprichnina. Probably the most skillful of Ivan's generals was Prince Alexander Borisovich Gorbaty-Shuysky, who advised the Tsar on military reform in the 1550s and presided over the Russian army during the siege and capture of Kazan in 1552. He was executed on fabricated charges in February 1565. Prince Ivan Petrovich Shuysky, also from a cadet line of the family, commanded the defence of Pskov during its prolonged siege by Stefan Báthory. Tsar Feodor, upon making Ivan Petrovich his military advisor, devolved on him enormous revenues supplied by Pskov's merchants. Soon enough, however, the Pskovian hero was found guilty of conspiring against Boris Godunov and exiled into Belozersk, where he died on November 16, 1588. The last of the Shuiskys The last of the Russian Shuyskys were four brothers - Vasily Ivanovich Shuysky (briefly Russian Tsar as Vasily IV), Dmitry Ivanovich Shuisky (infamous for having poisoned his brilliant cousin, Prince Mikhail Vasilyevich Skopin-Shuysky), Alexander Ivanovich Shuysky, and Ivan Ivanovich Shuysky "Pugovka" ("the Button"). All four were boyars and grandsons of Andrey Mikhailovich. The last scion of the family, Ivan Pugovka, was put in charge of the courts in Moscow during the reign of his brother-in-law Vasily IV. Pugovka outlived his brothers after he was taken with them into captivity