Phototrophs () are organisms that carry out photon capture to produce complex organic compounds (e.g. carbohydrates) and acquire energy. They use the energy from light to carry out various cellular metabolic processes. It is a common misconception that phototrophs are obligatorily photosynthetic. Many, but not all, phototrophs often photosynthesize: they anabolically convert carbon dioxide into organic material to be utilized structurally, functionally, or as a source for later catabolic processes (e.g. in the form of starches, sugars and fats). All phototrophs either use electron transport chains or direct proton pumping to establish an electrochemical gradient which is utilized by ATP synthase, to provide the molecular energy currency for the cell. Phototrophs can be either autotrophs or heterotrophs. If their electron and hydrogen donors are inorganic compounds (e.g. , as in some purple sulfur bacteria, or , as in some green sulfur bacteria) they can be also called lithotrophs, and so, some photoautotrophs are also called photolithoautotrophs. Examples of phototroph organisms are Rhodobacter capsulatus, Chromatium, and Chlorobium. History Originally used with a different meaning, the term took its current definition after Lwoff and collaborators (1946). Photoautotroph Most of the well-recognized phototrophs are autotrophic, also known as photoautotrophs, and can fix carbon. They can be contrasted with chemotrophs that obtain their energy by the oxidation of electron donors in their environments. Photoautotrophs are capable of synthesizing their own food from inorganic substances using light as an energy source. Green plants and photosynthetic bacteria are photoautotrophs. Photoautotrophic organisms are sometimes referred to as holophytic. Such organisms derive their energy for food synthesis from light and are capable of using carbon dioxide as their principal source of carbon. Oxygenic photosynthetic organisms use chlorophyll for light-energy capture and oxidize water, "splitting" it into molecular oxygen. In contrast, anoxygenic photosynthetic bacteria have a substance called bacteriochlorophyll – which absorbs predominantly at non-optical wavelengths – for light-energy capture, live in aquatic environments, and will, using light, oxidize chemical substances such as hydrogen sulfide rather than water. Ecology In an ecological context, phototrophs are often the food source for neighboring heterotrophic life. In terrestrial environments, plants are the predominant variety, while aquatic environments include a range of phototrophic organisms such as algae (e.g., kelp), other protists (such as euglena), phytoplankton, and bacteria (such as cyanobacteria). The depth to which sunlight or artificial light can penetrate into water, so that photosynthesis may occur, is known as the photic zone.