to most Pacific salmon, sockeye salmon are semelparous, meaning they die after spawning once. Some sockeye, called kokanee, do not migrate to the ocean and live their entire lives in freshwater lakes. The majority of sockeye spawn in rivers near lakes and juveniles will spend one to two years in the lake before migrating to the ocean, although some populations will migrate to saltwater in their first year. Adult sockeye will spend two to three years in the ocean before returning to freshwater. Females will spawn in 3–5 redds over a period of several days. The eggs usually hatch within six to nine weeks and the fry typically rear in lakes before migrating to the ocean. Reproduction Males partake in competitive and sneaking tactics, formation of hierarchies, and non-hierarchical groupings around females who are ready to mate. Reproductive success varies more in males than females. The greater variability in male reproduction is associated with the greater average size and exaggerated shape of males. Reproductive success in females is determined by the number of eggs she lays, her body size, and the survival of the eggs, which is due in part to the quality of the nest environment. Male spatial distribution depends on shifts in reproductive opportunities, physical traits of breeding sites, as well as the operational sex ratio (OSR) of the environment. Non-dominant males adopt a subordinate behavior, acting as a satellite to mated pairs. During spawning, a subordinate male will move quickly into the redd and release their sperm. Nearby dominant males from other redds will also do this. Male social status is positively correlated to length and dorsal hump size. Larger females tend to spawn in shallower water, which is preferred over deeper water. There is a dramatic sexual dimorphism at maturity. Males go through numerous morphological changes at maturation including, an increase in body depth, hump height, and snout length. Snout size also increases in females, but hump height and adipose fin length do not increase. This could mean that longer snout sizes are sexually selected, but hump height and adipose fin length are not. Females develop large gonads that are about 25% of the body mass. Females are responsible for parental care. They select, prepare, and defend a nest site until they die or are displaced. Males do not participate in parental care at all, and they move between females after egg deposition. Sexual selection and