and material from Sudan, late dynastic northern Egypt (Gizeh), Somalia, Asia and the Pacific Islands "show the Badarian series to be most similar to a series from the northeast quandrant of Africa and then to other Africans". Dental trait analysis of Badarian fossils conducted in a thesis study found that they were closely related to other Afroasiatic-speaking populations inhabiting Northeast Africa and the Maghreb. Among the ancient populations, the Badarians were nearest to other ancient Egyptians (Naqada, Hierakonpolis, Abydos and Kharga in Upper Egypt; Hawara in Lower Egypt), and C-Group and Pharaonic era skeletons excavated in Lower Nubia, followed by the A-Group culture bearers of Lower Nubia, the Kerma and Kush populations in Upper Nubia, the Meroitic, X-Group and Christian period inhabitants of Lower Nubia, and the Kellis population in the Dakhla Oasis. Among the recent groups, the Badari markers were morphologically closest to the Shawia and Kabyle Berber populations of Algeria as well as Bedouin groups in Morocco, Libya and Tunisia, followed by other Afroasiatic-speaking populations in the Horn of Africa. The Late Roman era Badarian skeletons from Kellis were also phenotypically distinct from those belonging to other populations in Sub-Saharan Africa. Naqada culture The Naqada culture is an archaeological culture of Chalcolithic Predynastic Egypt (c. 4000–3000 BC), named for the town of Naqada, Qena Governorate. It is divided in three sub-periods: Naqada I, II and III. A number of biological anthropological studies have found Naqada skeletal remains to have clear, African affinities. In 1996, Lovell and Prowse also reported the presence of individuals buried at Naqada in what they interpreted to be elite, high status tombs, showing them to be an endogamous ruling or elite segment of the local population at Naqada, which is more closely related to populations in northern Nubia than to neighbouring populations in southern Egypt. Specifically, they stated the Naqda samples were "more similar to the Lower Nubian protodynastic sample than they are to the geographically more proximate Egyptian samples" in Qena and Badari. Although, they found the skeletal samples from the Naqada cemeteries to be significantly different to protodynastic populations in northern Nubia and predynastic Egyptian samples from Badari and Qena, which were also significantly different to northern Nubian populations. Overall, both the elite and nonelite individuals in the Naqada cemeteries were more similar to each other than they were to the samples in northern Nubia or to samples from Badari and Qena