Warder Clyde Allee (June 5, 1885 – March 18, 1955) was an American ecologist. He is recognized to be one of the great pioneers of American ecology. As an accomplished zoologist and ecologist, Allee was best known and recognized for his research on social behavior, aggregations and distributions of animals in aquatic as well as terrestrial environments. Allee attended Earlham College and upon his graduation in 1908, pursued advanced studies at the University of Chicago where he received his PhD and graduated summa cum laude in 1912. Allee's most significant research occurred during his time at the University of Chicago and at the Marine Biological Laboratory at Woods Hole in Massachusetts. His research findings led to many publications, with the most notable being Principles of Animal Ecology and Animal Aggregations. Allee was married to author Marjorie Hill Allee and remained active in the field of biology until his death in 1955 at the age of 70. Early life Warder Clyde Allee was born on June 5, 1885, on a farm in Bloomingdale, Indiana to Mary Emily Newlin and John Wesley Allee. He is of French Huguenot heritage; the original surname was spelled d'Ailly. He was named after his paternal grandmother, Sarah Ann Warder. His father was orphaned as a child and grew up at the homes of various relatives in the Bloomingdale region. Clyde Allee attended a one-room country school and led his class in scholarship. At Bloomingdale Academy he was again at the head of his graduating class and the winner of the oratorical contest. At the age of seventeen, he taught country school for a year and then the fifth and sixth grades in the Bloomingdale elementary school for another year. Then, at nineteen, he began his first year at Earlham College. He was raised in the Society of Friends and married Marjorie Hill, whose Quaker ancestry extended back into the seventeenth century. His strong Quaker beliefs would play a large role in his research later in his academic and professional career. Personal life Family background combined with the Quaker influence at Earlham College contributed to the Quaker mold in which Allee was cast. In 1912, he married Marjorie Hill, a life-long member of the Society of Friends. Allee met Marjorie when she was a freshman at Earlham College while Allee was a senior. Marjorie was most interested in English Literature and Writing and would go on to