Marsh, who made light of the mistake. This led to antagonism between Cope, who was embarrassed by the mistake, and Marsh, who brought up the mistake repeatedly for decades. Marsh returned to the issue during their controversy in the New York Herald in the 1890s (Marsh claimed he had pointed out the error to Cope immediately), when their dispute gained widespread public attention. The argument was part of the "Bone Wars" rivalry between the two, and is well known in the history of paleontology. Because of Cope's reputation as a brilliant paleontologist, it has been questioned why he would make such an obvious anatomical error. It has been suggested that, as a unique specimen in 1868, the original Elasmosaurus may have been hard to interpret based on the knowledge available at the time. Also, Cope initially thought it consisted of two specimens of different animalsin an 1868 letter to LeConte, Cope had referred to the supposed "smaller specimen" as Discosaurus carinatus. Cope was only in his late twenties and not formally trained in paleontology, and may have been influenced by Leidy's mistake of reversing the vertebral column of Cimoliasaurus. In 2002 the American art historian Jane P. Davidson noted that the fact that other scientists early on had pointed out Leidy's error argues against this explanation, adding that Cope was not convinced he had made a mistake. Plesiosaur anatomy was sufficiently well known at the time that Cope should not have made the mistake, according to Davidson. Cope did little work on the specimen since his 1870 description, and it was kept in storage for nearly 30 years. It was only redescribed in detail in 2005 by the German paleontologist Sven Sachs. Known and possible fossil elements Today, the incomplete holotype specimen, cataloged as ANSP 10081, is the only definite specimen of Elasmosaurus. It was long exhibited, but is now stored in a cabinet with other assigned fragments. The specimen consists of the premaxillae, part of the hind-section of the right maxilla, two maxilla fragments with teeth, the front part of the dentaries, three more jaw fragments, two cranial fragments of indeterminable identity, 72 neck vertebrae, including the atlas and axis, 3pectoral vertebrae, 6back vertebrae, 4sacral vertebrae, 18 tail vertebrae, as well as rib fragments. In 2013 an incomplete neck vertebra centrum of the holotype that had been mentioned by Cope but thought to have been lost was rediscovered in