Multnomah village of similar name at the confluence. This type specimen was most likely the coastal cutthroat trout subspecies . Until the 1960s, populations of westslope cutthroat trout and Yellowstone cutthroat trout were lumped into one subspecies; Salmo clarkii lewisii. Biologists later split the group into two subspecies, christening the name westslope cutthroat trout with the lewisii name which honors explorer Meriwether Lewis and renaming the Yellowstone cutthroat trout Salmo bouvierii, the first name given to the Yellowstone cutthroat trout by David Starr Jordan in 1883 honoring a U.S. Army Captain Bouvier. In 1989, morphological and genetic studies indicated trout of the Pacific Basin were genetically closer to Pacific salmon (Oncorhynchus species) than to the Salmos–brown trout or Atlantic salmon of the Atlantic Basin. Thus, in 1989, taxonomic authorities moved the rainbow, cutthroat and other Pacific Basin trout into the genus Oncorhynchus. Subspecies Behnke in his salmon and trout handbook of 2002 recognized 14 subspecies of cutthroat trout that are each native to a separate geographic area. Not all of them were scientifically described, and different views on the taxonomic identities have been presented in some cases. It has been suggested that the cutthroat trout evolved from a common Oncorhynchus ancestor that migrated along the Pacific Coast and into the mountain west primarily via the Columbia and Snake River basins 3-5 million years ago, in the late Pliocene or early Pleistocene epochs. These epochs had repeated glacial and interglacial periods that would have caused repeated fracturing and isolation of cutthroat trout populations, eventually resulting in the different subspecies found today. The 14 subspecies are found in four evolutionary groups—Coastal, Westslope, Yellowstone and Lahontan. Description Throughout their native and introduced ranges, cutthroat trout vary widely in size, coloration and habitat selection. Their coloration can range from golden to gray to green on the back. Cutthroat trout can generally be distinguished from rainbow trout by the presence of basibranchial teeth at the base of tongue and a maxillary that extends beyond the posterior edge of the eye. Depending on subspecies, strain and habitat, most have distinctive red, pink, or orange linear marks along the underside of their mandibles in the lower folds of the gill plates. These markings are responsible for the common name "cutthroat", first given to the trout by outdoor writer Charles Hallock in an 1884 article in The American Angler. These markings are not unique to the species. Some