110,000 in 1944. Vilnius again grew in population by being the capital of the Lithuanian Soviet Socialist Republic (according to the 1989 census, it had 576,747 residents). Despite the fact that almost whole Lithuania suffered from a large emigration after the restoration of independence in 1990, the number of residents in Vilnius remained almost unchanged (542,287 in 2001) and began to steadily grow every year since 2006 to 580,020 residents (as of 1 January 2020). Historic ethnic makeup Around 1000 years AD, the confluence of the Neris and Vilnia rivers was densely populated by the Brushed Pottery culture, which had a half a hectare fortified settlement on the Gediminas' Hill. The tribes of this culture were common throughout present-day Lithuania, east of the Šventoji River and in western Belarus. The direct descendants of this culture were a Baltic tribe – the Aukštaitians (). According to a prominent researcher of Vilnius history Antanas Čaplinskas, who researched the surnames of Vilnius residents in the archive documents of the city, the oldest surviving surnames of Vilnius residents are Lithuanian. Pagan Lithuanians mostly lived at the northern foot of Gediminas' Hill and in the Crooked Castle. Later, following the invitation of Grand Duke Gediminas, merchants and craftsmen began to move to Vilnius from the cities of the German Hanseatic League, France, Italy and Spain, and replaced the Lithuanian surnames with German, Polish, and Russian surnames. In the late 14th century, during the reign of Grand Duke Algirdas, Vilnius already had a Ruthenian quarter () in the present-day Latako and Rusų Streets, as the trade relations between the Grand Duchy of Lithuania and the Ruthenian principalities were quite well developed, therefore quite a few Ruthenian merchants lived there and the Ruthenian nobles had their residences in the quarter. The variety of nations in Vilnius was further increased by Grand Duke Vytautas the Great, who introduced Litvak Jews, Tatars and Crimean Karaites. After a few hundred years, the number of locals in Vilnius was smaller than the number of newcomers. However, according to an analysis of the tax registers of 1572, Lithuania proper had 850,000 residents of which 680,000 were Lithuanians. Beginning during the Polish–Lithuanian Commonwealth, the Polish culture began to penetrate the city rapidly and soon the Polish language prevailed in the city, even the Magistrate's documents were written in Polish until the November Uprising in 1831. After living for a while in Vilnius, foreign