was used as a fixative for perfumes. The most successful whaling nations at this time were the Netherlands, Japan, and the United States. Commercial whaling was historically important as an industry well throughout the 17th, 18th and 19th centuries. Whaling was at that time a sizeable European industry with ships from Britain, France, Spain, Denmark, the Netherlands and Germany, sometimes collaborating to hunt whales in the Arctic, sometimes in competition leading even to war. By the early 1790s, whalers, namely the Americans and Australians, focused efforts in the South Pacific where they mainly hunted sperm whales and right whales, with catches of up to 39,000 right whales by Americans alone. By 1853, US profits reached US$11,000,000 (GB£6.5m), equivalent to US$348,000,000 (GB£230m) today, the most profitable year for the American whaling industry. Commonly exploited species included North Atlantic right whales, sperm whales, which were mainly hunted by Americans, bowhead whales, which were mainly hunted by the Dutch, common minke whales, blue whales, and grey whales. The scale of whale harvesting decreased substantially after 1982 when the International Whaling Commission (IWC) placed a moratorium which set a catch limit for each country, excluding aboriginal groups until 2004. Current whaling nations are Norway, Iceland, and Japan, despite their joining to the IWC, as well as the aboriginal communities of Siberia, Alaska, and northern Canada. Subsistence hunters typically use whale products for themselves and depend on them for survival. National and international authorities have given special treatment to aboriginal hunters since their methods of hunting are seen as less destructive and wasteful. This distinction is being questioned as these aboriginal groups are using more modern weaponry and mechanized transport to hunt with, and are selling whale products in the marketplace. Some anthropologists argue that the term "subsistence" should also apply to these cash-based exchanges as long as they take place within local production and consumption. In 1946, the IWC placed a moratorium, limiting the annual whale catch. Since then, yearly profits for these "subsistence" hunters have been close to US$31 million (GB£20m) per year. Other threats Whales can also be threatened by humans more indirectly. They are unintentionally caught in fishing nets by commercial fisheries as bycatch and accidentally swallow fishing hooks. Gillnetting and Seine netting is a significant cause of mortality in whales and other marine mammals. Species commonly entangled include beaked whales. Whales are also affected by marine pollution. High levels of