age. Additional bones of this individual (SHN (JJS)-65), including a left femur, a right tibia, and a partial left fibula (calf bone), were since exposed due to progressing cliff erosion. Although initially part of a private collection, these additional elements became officially curated after the private collection was donated to the Sociedade de História Natural in Torres Vedras, and were described in detail in 2015. The specimen was ascribed to the species Ceratosaurus dentisulcatus by Mateus and colleagues in 2006. A 2008 review by Carrano and Sampson confirmed the assignment to Ceratosaurus, but concluded that the assignment to any specific species is not possible at present. In 2015, Elisabete Malafaia and colleagues, who questioned the validity of C. dentisulcatus, assigned the specimen to Ceratosaurus aff. Ceratosaurus nasicornis. Other reports include a single tooth found in Moutier, Switzerland. Originally named by Janensch in 1920 as Labrosaurus meriani, the tooth was later assigned Ceratosaurus sp. (of unknown species) by Madsen and Welles. In 2008, Matías Soto and Daniel Perea described teeth from the Tacuarembó Formation in Uruguay, including a presumed premaxillary tooth crown. This shows vertical striations on its inner side and lacks denticles on its front edge; these features are, in this combination, only known from Ceratosaurus. The authors, however, stressed that an assignment to Ceratosaurus is infeasible as the remains are scant, and furthermore note that the assignment of the European and African material to Ceratosaurus has to be viewed with caution. In 2020, Soto and colleagues described additional Ceratosaurus teeth from the same formation that further support their earlier interpretation. Description Ceratosaurus followed the body plan typical for large theropod dinosaurs. A biped, it moved on powerful hind legs, while its arms were reduced in size. Specimen USNM 4735, the first discovered skeleton and holotype of Ceratosaurus nasicornis, was an individual or in length according to separate sources. Whether this animal was fully grown is not clear. Othniel Charles Marsh, in 1884, suggested that this specimen weighed about half as much as the contemporary Allosaurus. In more recent accounts, this was revised to , , or . Three additional skeletons discovered in the latter half of the 20th century were substantially larger. The first of these, UMNH VP 5278, was informally estimated by James Madsen to have been around long, but was later estimated at in length. Its weight was calculated at , , and in separate works. The