snout to dorsoventral bending compared with Spinosaurus and the gharial. The authors concluded (in contrast to the 2007 study) that Baryonyx performed differently than the gharial; spinosaurids were not exclusive piscivores, and their diet was determined by their individual size. In a 2014 conference abstract, the American palaeontologist Danny Anduza and Fowler pointed out that grizzly bears do not gaff fish out of the water as was suggested for Baryonyx, and also ruled out that the dinosaur would not have darted its head like herons, since the necks of spinosaurids were not strongly S-curved, and their eyes were not well-positioned for binocular vision. Instead, they suggested the jaws would have made sideways sweeps to catch fish, like the gharial, with the hand claws probably used to stamp down and impale large fish, whereafter they manipulated them with their jaws, in a manner similar to grizzly bears and fishing cats. They did not find the teeth of spinosaurids suitable for dismembering prey, due to their lack of serrations, and suggested they would have swallowed prey whole (while noting they could also have used their claws for dismemberment). A 2016 study by the Belgian palaeontologist Christophe Hendrickx and colleagues found that adult spinosaurs could displace their mandibular rami (halves of the lower jaw) sideways when the jaw was depressed, which allowed the pharynx (opening that connects the mouth to the oesophagus) to be widened. This jaw-articulation is similar to that seen in pterosaurs and living pelicans, and would likewise have allowed spinosaurids to swallow large prey such as fish and other animals. They also reported that the possible Portuguese Baryonyx fossils were found associated with isolated Iguanodon teeth, and listed it along with other such associations as support for opportunistic feeding behaviour in spinosaurs. Another 2016 study by the French palaeontologist Romain Vullo and colleagues found that the jaws of spinosaurids were convergent with those of pike conger eels; these fish also have jaws that are compressed side to side (whereas the jaws of crocodilians are compressed from top to bottom), an elongated snout with a "terminal rosette" that bears enlarged teeth, and a notch behind the rosette with smaller teeth. Such jaws likely evolved for grabbing prey in aquatic environments with low light, and may have helped in prey detection. Motion and aquatic habits In their original description, Charig and Milner did not consider Baryonyx to be aquatic (due to its