effects, posited in the theory, was false. Even though Waddington became critical of the neo-darwinian synthetic theory of evolution, he still described himself as a Darwinian, and called for an extended evolutionary synthesis based on his research. Reviewing the debate in 2015, the systems biologist Denis Noble writes however that As an organiser Waddington was very active in advancing biology as a discipline. He contributed to a book on the role of the sciences in times of war, and helped set up several professional bodies representing biology as a discipline. A remarkable number of his contemporary colleagues in Edinburgh became Fellows of the Royal Society during his time there, or shortly thereafter. Waddington was an old-fashioned intellectual who lived in both the arts and science milieus of the 1950s and wrote widely. His 1969 book Behind Appearance; a Study of the Relations Between Painting and the Natural Sciences in This Century (MIT press) not only has wonderful pictures but is still worth reading. Waddington was, without doubt, the most original and important thinker about developmental biology of the pre-molecular age and the medal of the British Society for Developmental Biology is named after him. Waddington co-founded The Institute for Advanced Studies in the Humanities at the University of Edinburgh in 1969 with Professor John MacQueen, Professor of Scottish Literature and Oral Tradition. Selected works Books Waddington, C. H. (1939). An Introduction to Modern Genetics. London : George Alien & Unwin. ––– (1940). Organisers & Genes. Cambridge: Cambridge University Press. ––– and others (1942). Science and Ethics, George Allen & Unwin. ––– (1946). How Animals Develop. London : George Allen & Unwin. ––– (1948). The Scientific Attitude, Pelican Books ––– (1953). The Epigenetics of birds. Cambridge : Cambridge University Press. ––– (1956). Principles of Embryology. London : George Allen & Unwin. ––– (1957). The Strategy of the Genes. London : George Allen & Unwin. ––– (1959). Biological Organisation Cellular and Subcellular : Proceedings of a Symposium. London: Pergamon Press. ––– (1960). The Ethical Animal. London : George Allen & Unwin. ––– (1961). The Human Evolutionary System. In: Michael Banton (Ed.), Darwinism and the Study of Society. London: Tavistock. ––– (1961). The Nature of Life. London : George, Allen, & Unwin. ––– (1962). New Patterns in Genetics and Development. New York: Columbia University Press. ––– (1966). Principles of Development and Differentiation. New York: Macmillan Company. ––– (1970). 72). Behind Appearance : A