the initial decades of its definition by the Pan-African Congress of Prehistory, it was parallel in Africa to the Upper and Middle Paleolithic. However, since then Radiocarbon dating has shown that the Middle Stone Age is in fact contemporaneous with the Middle Paleolithic. The Early Stone Age therefore is contemporaneous with the Lower Paleolithic and happens to include the same main technologies, Oldowan and Acheulean, which produced Mode 1 and Mode 2 stone tools respectively. A distinct regional term is warranted, however, by the location and chronology of the sites and the exact typology. Middle Stone Age (MSA) The Middle Stone Age was a period of African prehistory between Early Stone Age and Late Stone Age. It began around 300,000 years ago and ended around 50,000 years ago. It is considered as an equivalent of European Middle Paleolithic. It is associated with anatomically modern or almost modern Homo sapiens. Early physical evidence comes from Omo and Herto, both in Ethiopia and dated respectively at c. 195 ka and at c. 160 ka. Later Stone Age (LSA) The Later Stone Age (LSA, sometimes also called the Late Stone Age) refers to a period in African prehistory. Its beginnings are roughly contemporaneous with the European Upper Paleolithic. It lasts until historical times and this includes cultures corresponding to Mesolithic and Neolithic in other regions. Material culture Tools Stone tools were made from a variety of stones. For example, flint and chert were shaped (or chipped) for use as cutting tools and weapons, while basalt and sandstone were used for ground stone tools, such as quern-stones. Wood, bone, shell, antler (deer) and other materials were widely used, as well. During the most recent part of the period, sediments (such as clay) were used to make pottery. Agriculture was developed and certain animals were domesticated as well. Some species of non-primates are able to use stone tools, such as the sea otter, which breaks abalone shells with them. Primates can both use and manufacture stone tools. This combination of abilities is more marked in apes and men, but only men, or more generally Hominans, depend on tool use for survival. The key anatomical and behavioral features required for tool manufacture, which are possessed only by Hominans, are the larger thumb and the ability to hold by means of an assortment of grips. Food and drink Food sources of the Palaeolithic hunter-gatherers were wild plants