types of inheritance: Mendelian inheritance of variation within species, and Non-Mendelian inheritance of variation in a macroevolutionary sense. Dobzhansky later stated that Filipchenko “bet on the wrong horse”. He followed Morgan to the California Institute of Technology from 1930 to 1940. On the basis of his experiments, he articulated the idea that reproductive isolation can be caused by differences in presence of microbial symbionts between populations. In 1937, he published one of the major works of the modern evolutionary synthesis, the synthesis of evolutionary biology with genetics, titled Genetics and the Origin of Species, which amongst other things, defined evolution as "a change in the frequency of an allele within a gene pool". Dobzhansky's work was instrumental in spreading the idea that it is through mutations in genes that natural selection takes place. Also in 1937, he became a naturalized citizen of the United States. During this time, he had a very public falling out with one of his Drosophila collaborators, Alfred Sturtevant, based primarily in professional competition. He returned to Columbia University from 1940 to 1962. Among his students was geneticist Bruce Wallace. In 1941, Dobzhansky was awarded the Daniel Giraud Elliot Medal from the National Academy of Sciences. In 1943, the University of São Paulo awarded him an honorary doctorate. He was one of the signatories of the 1950 UNESCO statement The Race Question. He then moved to the Rockefeller Institute (shortly to become Rockefeller University) until his retirement in 1971. In 1972 he was elected the founding president of the Behavior Genetics Association, and was recognized by the society for his role in behavior genetics, and the founding of the society by the creation of the Dobzhansky Award (for a lifetime of outstanding scholarship in behavior genetics). Dobzhansky’s work in the field of evolutionary genetics, with the help of Sewall Wright, integrated standards of the theoretical, natural historical, and experimental work. Dobzhansky was elected a Foreign Member of the Royal Society (ForMemRS) in 1965. In 1970, he published Genetics of the evolutionary process. Dobzhansky was renowned as the president of the Genetics Society of America in 1941, president of the American Society of Naturalists in 1950, president of the Society for the Study of Evolution in 1951, president of the American Society of Zoologists in 1963, a member of the Board of Directors of the American Eugenics Society in 1964, and president of the American Teilhard de