Early use and development Herodotus mentions that the Egyptian pharaoh Necho II (610–595 BC) built triremes on the Nile, for service in the Mediterranean, and in the Red Sea, but this reference is disputed by modern historians, and attributed to a confusion, since "triērēs" was by the 5th century used in the generic sense of "warship", regardless its type. The first definite reference to the use of triremes in naval combat dates to ca. 525 BC, when, according to Herodotus, the tyrant Polycrates of Samos was able to contribute 40 triremes to a Persian invasion of Egypt (Battle of Pelusium). Thucydides meanwhile clearly states that in the time of the Persian Wars, the majority of the Greek navies consisted of (probably two-tiered) penteconters and ploia makrá ("long ships"). In any case, by the early 5th century, the trireme was becoming the dominant warship type of the eastern Mediterranean, with minor differences between the "Greek" and "Phoenician" types, as literary references and depictions of the ships on coins make clear. The first large-scale naval battle where triremes participated was the Battle of Lade during the Ionian Revolt, where the combined fleets of the Greek Ionian cities were defeated by the Persian fleet, composed of squadrons from their Phoenician, Carian, and Egyptian subjects. The Persian Wars Athens was at that time embroiled in a conflict with the neighbouring island of Aegina, which possessed a formidable navy. In order to counter this, and possibly with an eye already at the mounting Persian preparations, in 483/2 BC the Athenian statesman Themistocles used his political skills and influence to persuade the Athenian assembly to start the construction of 200 triremes, using the income of the newly discovered silver mines at Laurion. The first clash with the Persian navy was at the Battle of Artemisium, where both sides suffered great casualties. However, the decisive naval clash occurred at Salamis, where Xerxes' invasion fleet was decisively defeated. After Salamis and another Greek victory over the Persian fleet at Mycale, the Ionian cities were freed, and the Delian League was formed under the aegis of Athens. Gradually, the predominance of Athens turned the League effectively into an Athenian Empire. The source and foundation of Athens' power was her strong fleet, composed of over 200 triremes. It not only secured control of the Aegean Sea and the loyalty of her allies, but also safeguarded the trade routes and the