institution to have a tuatara breeding programme; starting in 1986 they bred S. punctatus and have focused on S. guntheri more recently. Hamilton Zoo, Auckland Zoo and Wellington Zoo also breed tuatara for release into the wild. At Auckland Zoo in the 1990s it was discovered that tuatara have temperature-dependent sex determination. The Victoria University of Wellington maintains a research programme into the captive breeding of tuatara, and the Pukaha / Mount Bruce National Wildlife Centre keeps a pair and a juvenile. The WildNZ Trust has a tuatara breeding enclosure at Ruawai. One notable captive breeding success story took place in January 2009, when all 11 eggs belonging to 110 year-old tuatara Henry and 80 year-old tuatara Mildred hatched. This story is especially remarkable as Henry required surgery to remove a cancerous tumour in order to successfully breed. In January 2016, Chester Zoo, England, announced that they succeeded in breeding the tuatara in captivity for the first time outside its homeland. Cultural significance Tuatara feature in a number of indigenous legends, and are held as ariki (God forms). Tuatara are regarded as the messengers of Whiro, the god of death and disaster, and Māori women are forbidden to eat them. Tuatara also indicate tapu (the borders of what is sacred and restricted), beyond which there is mana, meaning there could be serious consequences if that boundary is crossed. Māori women would sometimes tattoo images of lizards, some of which may represent tuatara, near their genitals. Today, tuatara are regarded as a taonga (special treasure) along with being viewed as the kaitiaki (guardian) of knowledge. The tuatara was featured on one side of the New Zealand five-cent coin, which was phased out in October 2006. Tuatara was also the name of the Journal of the Biological Society of Victoria University College and subsequently Victoria University of Wellington, published from 1947 until 1993. It has now been digitised by the New Zealand Electronic Text Centre, also at Victoria. In popular culture A tuatara named "Tua" is prominently featured in the 2017 novel Turtles All the Way Down by John Green. The tuatara was the inspiration for a DC Comics superhero, also with a third eye, called Tuatara, member of the Global Guardians. There is a brand of New Zealand craft beer named after the Tuatara which particularly references the third eye in its advertising. The Tuatara hypercar, designed and manufactured by SSC