there stands of white spruce older than 250 years. The prevalence of fire-adaptive morphologic and reproductive characteristics of many boreal plant species is further evidence pointing to a long and intimate association with fire. Seven of the ten most common trees in the boreal forest—jack pine, lodgepole pine, aspen, balsam poplar (Populus balsamifera), paper birch, tamarack, black spruce – can be classed as pioneers in their adaptations for rapid invasion of open areas. White spruce shows some pioneering abilities, too, but is less able than black spruce and the pines to disperse seed at all seasons. Only balsam fir and alpine fir seem to be poorly adapted to reproduce after fire, as their cones disintegrate at maturity, leaving no seed in the crowns. The oldest forests in the northwest boreal region, some older than 300 years, are of white spruce occurring as pure stands on moist floodplains. Here, the frequency of fire is much less than on adjacent uplands dominated by pine, black spruce and aspen. In contrast, in the Cordilleran region, fire is most frequent in the valley bottoms, decreasing upward, as shown by a mosaic of young pioneer pine and broadleaf stands below, and older spruce–fir on the slopes above. Without fire, the boreal forest would become more and more homogeneous, with the long-lived white spruce gradually replacing pine, aspen, balsam poplar, and birch, and perhaps even black spruce, except on the peatlands. Climate change During the last quarter of the twentieth century, the zone of latitude occupied by the boreal forest experienced some of the greatest temperature increases on Earth. Winter temperatures have increased more than summer temperatures. In summer, the daily low temperature has increased more than the daily high temperature. The number of days with extremely cold temperatures (e.g., −20 to −40 °C (−4 to −40 °F) has decreased irregularly but systematically in nearly all the boreal region, allowing better survival for tree-damaging insects. In Fairbanks, Alaska, the length of the frost-free season has increased from 60 to 90 days in the early twentieth century to about 120 days a century later. It has been hypothesized that the boreal environments have only a few states which are stable in the long term - a treeless tundra/steppe, a forest with >75% tree cover and an open woodland with ∼20% and ∼45% tree cover. Thus, continued climate change would be able to force at least some of