ecological characteristics, fishing intensity and population size. In the Coral Triangle, species at lower trophic levels that have smaller maximum sizes, faster growth and maturation rates and shorter life spans tend to recover more quickly than species having the opposite characteristics. For example, in the Philippines, populations of planktivores (e.g., fusiliers) and some herbivores (e.g., parrotfishes) recovered in <5–10 years in marine reserves, while predators (e.g., groupers) took 20–40 years. Increased fishing pressure adversely affects recovery rates (e.g., Great Barrier Reef and Papua New Guinea). Long-term protection allows species with slower recovery rates to achieve and maintain ecosystem health and associated fishery benefits. Permanent protection protects these species over the long-term. Short-term protections do not allow slow-recovering species to reach or maintain stable populations. In some Coral Triangle countries (e.g., Papua New Guinea and Solomon Islands), short term protections are the most common form of traditional marine resource management. These protections can help address problems at lower trophic levels (e.g., herbivores) or allow spawning to succeed. Other reasons for adopting short-term protections include allowing communities to stockpile resources for feasts or close areas for cultural reasons. Short-term/periodic reserves also may function as partial insurance by enhancing overall ecosystem resilience against catastrophes. Reopened reserves can be protected by management controls that limit the harvest to less than the increase achieved during closure, although at greatly reduced recovery rates. Resilience Some habitats and species are better prepared environmental changes or extremes. These include coral communities that handle high sea surface temperature (SST); areas with variable SSTs and carbonate chemistry and areas adjacent to undeveloped low-lying inland areas that coastal habitats can expand into as sea levels rise. Such areas constitute climate change refugia and can potentially better protect biodiversity than more fragile areas. They may also provide fishery benefits, since habitat loss from climate change is a major fishery threat. Local practices Local practices such as overfishing, blast fishing, trawling, coastal development and pollution threaten many marine habitats. These threats decrease ecosystem health and productivity and adversely affect focal and other species. Such practices can also decrease resilience. Some practices that originate beyond reserve boundaries (e.g., runoff) can be mitigated by considering their impacts within broader management frameworks. Areas that are not threatened by such practices and that are adjacent to other unthreatening areas may be better choices for reserves. Networks Networks of marine reserves can support both fisheries management and biodiversity