(e.g., an offshore reef) can reduce exports where that is desired. Habitats Minimum sustainable population sizes have not been determined for most marine populations. Instead, fisheries ecologists use the fraction of unfished stock levels as a proxy. Meta-analyses suggest that maintaining populations above ∼37% of those levels generally ensures stable populations, although variations in fishing pressure allow fractions as small as 10% or as large as 40% (to protect species such as sharks and some grouper that have lower reproductive output or slower maturation). Higher fractions of habitat protection may protect areas vulnerable to disturbances such as typhoons or climate change. 20–30% protection can achieve fisheries objectives in areas with controlled fishing pressure and is the minimum level of habitat protection recommended by IUCN-WCPA. Special areas Many fish species congregate to facilitate spawning. Such congregations are spatially and temporally predictable and increase the species' vulnerability to overfishing. Species such as groupers and rabbitfishes travel long distances to congregate for days or weeks. Such gatherings are their only opportunities to reproduce and are crucial to population maintenance. Species such as snappers and parrotfishes congregate in feeding or resting areas. Juveniles may congregate in nursery areas without adults. Such special areas may require only seasonal protections if at other times no vital activities are taking place. Such reserves must be spaced to allow focal species to journey among them. If the location of such special areas is unknown, or is too large to include in a reserve, management approaches such as seasonal capture and sales restrictions may provide some protection. Sea turtle nesting areas, dugong feeding areas, cetacean migratory corridors and calving grounds are examples of other special areas that can be protected seasonally. Other types of special areas include isolated habitats that have unique assemblages and populations, habitats that are important for endemic species and highly diverse areas. Isolated populations Isolated populations (e.g.,those on remote atolls) have high conservation value where they harbor endemic species and/or unique assemblages. A location or population 20–30 km from its nearest neighbor generally qualifies as isolated in the absence of a persistent linking current. Their isolation (low connectivity) requires such areas to be largely self-replenishing. This leaves them less resilient to disturbance. Sustaining their marine species requires a higher fraction of living areas to be protected. Recovery Coral reef fish species recovery rates (from e.g., overfishing) depend on their life history and factors such as