It reached the same conclusion: the original broadly circumscribed Tibouchina consisted of four monophyletic clades. The authors proposed a split into four genera: a more narrowly circumscribed Tibouchina, two re-established genera Pleroma and Chaetogastra, and a new genus, Andesanthus. The relationship between Chaetogastra and the genus Brachyotum differed between a maximum likelihood analysis and a Bayesian inference analysis: the former found Brachyotum embedded within Chaetogastra, the latter found the two to be sisters. The part of their maximum likelihood cladogram which includes former Tibouchina species is as follows, using their genus names and with shading added to show the original broadly circumscribed Tibouchina s.l.: As re-circumscribed, Tibouchina is monophyletic and contains species belonging to the traditional sections T. section Tibouchina and T. section Barbigerae. Diagnostic characteristics include the presence of scale-like trichomes on the hypanthium and leaves and a long pedoconnective on lilac anthers, and the absence of glandular trichomes. Species are found in savanna habitats. Species , Plants of the World Online accepts the following species within Tibouchina: Tibouchina aegopogon (Naudin) Cogn. Tibouchina albescens Cogn. ex P.J.F.Guim., A.L.F.Oliveira & R.Romero Tibouchina alpestris Cogn. Tibouchina araguaiensis P.J.F.Guim. Tibouchina aspera Aubl. Tibouchina barbigera (Naudin) Baill. Tibouchina bicolor (Naudin) Cogn. Tibouchina bipenicillata (Naudin) Cogn. Tibouchina brevisepala Cogn. Tibouchina bruniana P.J.F.Guim. Tibouchina caatingae J.G.Freitas Tibouchina calycina Cogn. Tibouchina catharinae Pittier Tibouchina cujabensis Cogn. Tibouchina dissitiflora Wurdack Tibouchina duidae Gleason Tibouchina edmundoi Brade Tibouchina exasperata (Naudin) Cogn. Tibouchina fraterna N.E.Br. Tibouchina huberi Wurdack Tibouchina itatiaiae Cogn. Tibouchina johnwurdackiana Todzia Tibouchina karstenii Cogn. Tibouchina lithophila Wurdack Tibouchina llanorum Wurdack Tibouchina mathaei Cogn. Tibouchina melastomoides (Naudin) Cogn. Tibouchina nigricans Cogn. ex P.J.F.Guim., A.L.F.Oliveira & R.Romero Tibouchina octopetala Cogn. Tibouchina papyrus (Pohl) Toledo Tibouchina pogonanthera (Naudin) Cogn. Tibouchina rosanae P.J.F.Guim. & Woodgyer Tibouchina sickii Brade Tibouchina sipapoana Gleason Tibouchina spruceana Cogn. Tibouchina steyermarkii Wurdack Tibouchina striphnocalyx (DC.) Pittier Tibouchina verticillaris Cogn. Tibouchina xochiatencana de Santiago Selected former species Species placed in Tibouchina in its former broad sense include: Tibouchina anderssonii Wurdack, synonym of Chaetogastra anderssonii Tibouchina asperior (Cham.) Cogn., synonym of Pleroma asperius Tibouchina campii Wurdack, synonym of Chaetogastra campii Tibouchina ciliaris (Vent.) Cogn., synonym of Chaetogastra ciliaris Tibouchina clinopodifolia (DC.) Cogn., synonym of Chaetogastra clinopodifolia Tibouchina elegans Cogn., synonym of Pleroma elegans Tibouchina francavillana Cogn., synonym of Pleroma francavillanum Tibouchina gleasoniana Wurdack, synonym of Andesanthus gleasonianus Tibouchina granulosa, synonym of Pleroma granulosum Tibouchina gracilis (Bonpl.) Cogn., synonym of Chaetogastra gracilis Tibouchina grossa, synonym of Chaetogastra grossa