he was elected a Foreign Honorary Member of the American Academy of Arts and Sciences. He received one of the thirteen Darwin-Wallace Medals awarded by the Linnean Society of London in 2008; at that time the award was made only every 50 years. He was awarded the Darwin Medal of the Royal Society in 2010 'for his original and influential contributions to our understanding of the genetic basis of evolution'. In 1959 he published Berber Village, an account of an Oxford University expedition to the High Atlas mountains of Morocco. References 1932 births Academics of the University of Edinburgh Academics of the University of Nottingham Alumni of Magdalen College, Oxford British biologists Fellows of the Royal Society British geneticists 2014 deaths Fellows of the American Academy of Arts and Sciences Members of the American Philosophical Society