males being only slightly larger than the females. The average weight is . During the summer, the white-tailed ptarmigan is a speckled grayish brown with white underparts, tail and wings. In the fall, the plumage has turned a much more reddish-brown color and white feathers begin to grow through. By winter all the summer brown feathers are lost and the bird is completely white. A further molt in the spring precedes the breeding season and the bird returns to its summer plumage. The finely barred greyish coloration on the back makes it easy to distinguish this species from the much browner willow ptarmigan and rock ptarmigan. Both sexes maintain white tail and wing feathers all the year and males can be identified by their reddish eyecombs (fleshy growths above the eye), also present year-long. In general this bird is silent but it sometimes makes quiet, low-pitched hoots and soft clucking noises. Distribution The white-tailed ptarmigan is an alpine species, a permanent resident of the high mountains above or near the timber line. It occupies open country and flies a great deal more than forest grouse, but still prefers running to flying. It ranges from Alaska and western Canada south to northern New Mexico. Males return from their wintering areas to establish territories on spruce-willow timber line breeding grounds in April. Females arrive in early May and pairs are formed. The white-tailed ptarmigan is the only bird in North America to reside permanently in the alpine zone. Its habitat includes areas of boulders, krummholz, snowfields, rock slides, frost-heaved soil and upland herbage. Even in winter it stays in high valleys and mountain slopes where alder, willow, birch and spruce poke through the snow cover. The white-tailed ptarmigan was introduced into the Sierra Nevada of California and Uinta Mountains of Utah in the 1970s. It may have been native here during the early Pleistocene but became locally extinct due to climate changes with greater snow-cover in spring impacting on its breeding season. Alternatively, it may have been unable to colonize the Sierra Nevada because of the barriers provided by the Columbia River and the Great Basin, and the low altitudes of the intervening South Cascades. The white-tailed ptarmigan was first introduced into Oregon's Wallowa Mountains with 36 birds taken from Colorado and Washington in September 1967. A second set of 54 birds was released in September 1968 that were supplied from Colorado