at 1.0 m/s or above, which matches human walking speeds. The results of other studies have also supported the theory of a human-like gait. Hominina footprints from site S In 2015, footprints of the same age as the first reported footprints were unearthed at a site approximately 150 meters south of the original site G footprints. This site is called site S, and the 2 individuals who made the prints are named S1 and S2. S2 is represented by only 1 print, but S1 left a track of prints, the first 4 of which are shown in the composite image, along with an analysis of step and stride lengths. Further analysis indicated that individual S1 was considerably larger than any of the three individuals from site G. Other footprints and artifacts Other prints show the presence of twenty different animal species besides the hominin A. afarensis, among them hyenas, wild cats (Machairodus), baboons, wild boars, giraffes, gazelles, rhinos, several kinds of antelope, Hipparion, buffaloes, elephant relatives (of the extinct genus Deinotherium), hares and birds. Rain-prints can be seen as well. Few footprints are superimposed, which indicates that they were rapidly covered up. Most of the animals are represented by skeletal remains discovered in the area. No artifacts have been found in the vicinity, at least within the ancient Laetolil Beds that contain the trackway. However, artifacts from the younger Olpiro and Ngaloba Beds, also preserved at Laetoli, have been found. Interpretation and significance Before the discovery of the Laetoli footprints, there was much debate as to which developed first in the human evolutionary time line: a larger brain or bipedalism. The discovery of these footprints settled the issue, proving that the Laetoli hominins were fully bipedal long before the evolution of the modern human brain, and were bipedal close to a million years before the earliest known stone tools were made. The footprints were classified as possibly belonging to Australopithecus afarensis. Some analysts have noted in their interpretations that the smaller trail bears "telltale signs that suggest whoever left the prints was burdened on one side." This may suggest that a female was carrying an infant on her hip but this cannot be proven for certain. The footprints themselves were an unlikely discovery because they closely resemble modern human footprints, despite being almost 4 million years old. It is noted that the toe pattern is much the same as the