first assembled at sea in October 2020 as part of a rehearsal for its first operational deployment in 2021. In 2019, the Royal Navy announced the formation of two Littoral Response Groups as part of a transformation of its amphibious forces. These forward-based special operations-capable task groups are to be rapidly-deployable and able to carry out a range of tasks within the littoral, including raids and precision strikes. The first one, based in Europe, became operational in 2021, whilst the second will be based in the Indo-Pacific from 2023. They will centre around two amphibious assault ships, a company of Royal Marines and supporting elements. Command, control and organisation The titular head of the Royal Navy is the Lord High Admiral, a position which was held by the Duke of Edinburgh from 2011 until his death in 2021 and since then remains vacant. The position had been held by Queen Elizabeth II from 1964 to 2011; the Sovereign is the Commander-in-chief of the British Armed Forces. The professional head of the Naval Service is the First Sea Lord, an admiral and member of the Defence Council of the United Kingdom. The Defence Council delegates management of the Naval Service to the Admiralty Board, chaired by the Secretary of State for Defence, which directs the Navy Board, a sub-committee of the Admiralty Board comprising only naval officers and Ministry of Defence (MOD) civil servants. These are all based in MOD Main Building in London, where the First Sea Lord, also known as the Chief of the Naval Staff, is supported by the Naval Staff Department. Organisation The Fleet Commander has responsibility for the provision of ships, submarines and aircraft ready for any operations that the Government requires. Fleet Commander exercises his authority through the Navy Command Headquarters, based at in Portsmouth. An operational headquarters, the Northwood Headquarters, at Northwood, London, is co-located with the Permanent Joint Headquarters of the United Kingdom's armed forces, and a NATO Regional Command, Allied Maritime Command. The Royal Navy was the first of the three armed forces to combine the personnel and training command, under the Principal Personnel Officer, with the operational and policy command, combining the Headquarters of the Commander-in-Chief, Fleet and Naval Home Command into a single organisation, Fleet Command, in 2005 and becoming Navy Command in 2008. Within the combined command, the Second Sea Lord continues to act as the Principal Personnel Officer.