Pterodactylus (from Greek () meaning 'winged finger') is an extinct genus of pterosaurs. It is thought to contain only a single species, Pterodactylus antiquus, which was the first pterosaur to be named and identified as a flying reptile. Fossil remains of Pterodactylus have primarily been found in the Solnhofen limestone of Bavaria, Germany, which dates from the Late Jurassic period (early Tithonian stage), about 150.8 to 148.5 million years ago. More fragmentary remains of Pterodactylus have tentatively been identified from elsewhere in Europe and in Africa. Pterodactylus was a generalist carnivore that probably fed on a variety of invertebrates and vertebrates. Like all pterosaurs, Pterodactylus had wings formed by a skin and muscle membrane stretching from its elongated fourth finger to its hind limbs. It was supported internally by collagen fibres and externally by keratinous ridges. Pterodactylus was a small pterosaur compared to other famous genera such as Pteranodon and Quetzalcoatlus, and it also lived earlier, during the Late Jurassic period, while both Pteranodon and Quetzalcoatlus lived during the Late Cretaceous. Pterodactylus lived alongside other small pterosaurs such as the well-known Rhamphorhynchus, as well as other genera such as Scaphognathus, Anurognathus and Ctenochasma. Pterodactylus is classified as an early-branching member of the ctenochasmatid lineage, within the pterosaur clade Pterodactyloidea. Discovery and history The type specimen of the animal now known as Pterodactylus antiquus was the first pterosaur fossil ever to be identified. The first Pterodactylus specimen was described by the Italian scientist in 1784, based on a fossil skeleton that had been unearthed from the Solnhofen limestone of Bavaria. Collini was the curator of the , or nature cabinet of curiosities (a precursor to the modern concept of the natural history museum), in the palace of Charles Theodore, Elector of Bavaria at Mannheim. The specimen had been given to the collection by Count around 1780, having been recovered from a lithographic limestone quarry in . The actual date of the specimen's discovery and entry into the collection is unknown however, and it was not mentioned in a catalogue of the collection taken in 1767, so it must have been acquired at some point between that date and its 1784 description by Collini. This makes it potentially the earliest documented pterosaur find; the "Pester Exemplar" of the genus Aurorazhdarcho was described in 1779 and possibly discovered earlier than the Mannheim specimen, but it was at first considered to be a fossilized