or taking bribes outright. Quaestors' behaviour did not always comport with their administrative and legal responsibilities. On campaign On campaign, provincial quaestors acted as subordinate military officers to their attached superior, taking a role "analogous to... that [of] other members of the governor's entourage, such as his legates". At times, the quaestor could get into tension with the governor's legates over respective spheres of responsibility or accountability; officially, however, the quaestor was "higher up the chain of command... [as], besides the governor, he was the only magistrate [and] representative of the Senate and the Roman people", giving him "greater authority than legates in all areas of provincial command". Quaestors are documented at various times leading and raising troops and fleets under the command of their governors. Some quaestors were delegated significant open-ended responsibilities far exceeding administrative tasks: Lucullus, for example, during the First Mithridatic War as Sulla's proquaestor, led troops, assembled fleets, travelled the eastern Mediterranean as a diplomat, intervened to overthrow governments, commanded naval battles, captured prisoners, and levied taxes and indemnities. When a governor left the province, he normally left it to his quaestor's command (though this was at times given instead to one of his high-ranking legates). If a governor died, however, the quaestor generally assumed command of the forces until replacement, possibly with imperium pro praetore. The specifics of how this imperium was delegated after the death of its actual possessor are unclear: some scholars believe that this was automatic, whereas others believed that a proconsul had to first endow his quaestor with propraetorian imperium. A provincial quaestor also could be sent as a diplomatic representative. Two famous examples thereof are those of Tiberius Gracchus and Sulla: Gracchus negotiated a peace treaty on behalf of his proconsul allowing some twenty thousand soldiers to leave with their lives (though the treaty was later invalidated by the Senate) and Sulla negotiated the capture of Jugurtha at the end of the Jugurthine War. History There were initially two quaestors; they were initially appointed by the consuls, but after 447 BC, they were elected by the comitia tributa. When plebeians were permitted to stand for the quaestorship in 421 BC, two more were added, with assignments to administer the aerarium under senatorial direction. It is also around this time that Livy reports a relationship between the quaestors and the public treasury. After 267 BC, four more quaestors were added, possibly