cordaitaleans occurred approximately 252.76 Ma, around 820,000 years before the end-Permian extinction in South China, suggesting that the end-Permian biotic catastrophe may have started earlier on land and that the ecological crisis may have been more gradual and asynchronous on land compared to its more abrupt onset in the marine realm. In South China, the subtropical Cathaysian gigantopterid dominated rainforests abruptly collapsed. The floral extinction in South China is associated with bacterial blooms in soil and nearby lacustrine ecosystems, with soil erosion resulting from the die-off of plants being their likely cause. Terrestrial vertebrates There is enough evidence to indicate that over two thirds of terrestrial labyrinthodont amphibians, sauropsid ("reptile") and therapsid ("proto-mammal") taxa became extinct. Large herbivores suffered the heaviest losses. All Permian anapsid reptiles died out except the procolophonids (although testudines have morphologically-anapsid skulls, they are now thought to have separately evolved from diapsid ancestors). Pelycosaurs died out before the end of the Permian. Too few Permian diapsid fossils have been found to support any conclusion about the effect of the Permian extinction on diapsids (the "reptile" group from which lizards, snakes, crocodilians, and dinosaurs (including birds) evolved). The groups that survived suffered extremely heavy losses of species and some terrestrial vertebrate groups very nearly became extinct at the end of the Permian. Some of the surviving groups did not persist for long past this period, but others that barely survived went on to produce diverse and long-lasting lineages. However, it took 30million years for the terrestrial vertebrate fauna to fully recover both numerically and ecologically. It is difficult to analyze extinction and survival rates of land organisms in detail because few terrestrial fossil beds span the Permian–Triassic boundary. The best-known record of vertebrate changes across the Permian–Triassic boundary occurs in the Karoo Supergroup of South Africa, but statistical analyses have so far not produced clear conclusions. Biotic recovery In the wake of the extinction event, the ecological structure of present-day biosphere evolved from the stock of surviving taxa. In the sea, the "Modern Evolutionary Fauna" became dominant over elements of the "Palaeozoic Evolutionary Fauna". Typical taxa of shelly benthic faunas were now bivalves, snails, sea urchins and Malacostraca, whereas bony fishes and marine reptiles diversified in the pelagic zone. On land, dinosaurs and mammals arose in the course of the Triassic. The profound change in the taxonomic composition was partly a result of the selectivity of the