days of life, as in modern reptiles, rather than depend on parents for food. Fossilised Hamipterus nests were shown preserving many male and female pterosaurs together with their eggs in a manner to a similar to that of modern seabird colonies. Due to how underdeveloped the chests of the hatchlings were for flying, it was suggested that Hamipterus may have practiced some form of parental care. However, this study has since been criticised. Most evidence currently leans towards pterosaur hatchlings being superprecocial, similar to that of megapode birds, which fly after hatching without the need of parental care. A further study compares evidence for superprecociality and "late term flight" and overwhelmingly suggests that most if not all pterosaurs were capable of flight soon after hatching. Growth rates of pterosaurs once they hatched varied across different groups. In more primitive, long-tailed pterosaurs ("rhamphorhynchoids"), such as Rhamphorhynchus, the average growth rate during the first year of life was 130% to 173%, slightly faster than the growth rate of alligators. Growth in these species slowed after sexual maturity, and it would have taken more than three years for Rhamphorhynchus to attain maximum size. In contrast, the more advanced, large pterodactyloid pterosaurs, such as Pteranodon, grew to adult size within the first year of life. Additionally, pterodactyloids had determinate growth, meaning that the animals reached a fixed maximum adult size and stopped growing. A 2021 study indicates that pterosaur juveniles of larger species increasingly took the roles previously occupied by adult small pterosaurs. Daily activity patterns Comparisons between the scleral rings of pterosaurs and modern birds and reptiles have been used to infer daily activity patterns of pterosaurs. The pterosaur genera Pterodactylus, Scaphognathus, and Tupuxuara have been inferred to be diurnal, Ctenochasma, Pterodaustro, and Rhamphorhynchus have been inferred to be nocturnal, and Tapejara has been inferred to be cathemeral, being active throughout the day for short intervals. As a result, the possibly fish-eating Ctenochasma and Rhamphorhynchus may have had similar activity patterns to modern nocturnal seabirds, and the filter-feeding Pterodaustro may have had similar activity patterns to modern anseriform birds that feed at night. The differences between activity patterns of the Solnhofen pterosaurs Ctenochasma, Rhamphorhynchus, Scaphognathus, and Pterodactylus may also indicate niche partitioning between these genera. Cultural significance Pterosaurs have been a staple of popular culture for as long as their cousins the dinosaurs, though they are usually not featured as prominently in films,