also confirmed substantial Balkan and Middle Eastern ancestry. A study from 2001 by Gresham et al. suggests "a limited number of related founders, compatible with a small group of migrants splitting from a distinct caste or tribal group". The same study found that "a single lineage... found across Romani populations, accounts for almost one-third of Romani males". A 2004 study by Morar et al. concluded that the Romani population "was founded approximately 32–40 generations ago, with secondary and tertiary founder events occurring approximately 16–25 generations ago". Haplogroup H-M82 is a major lineage cluster in the Balkan Romani group, accounting for approximately 60% of the total. Haplogroup H is uncommon in Europe but present in the Indian subcontinent and Sri Lanka. A study of 444 people representing three ethnic groups in North Macedonia found mtDNA haplogroups M5a1 and H7a1a were dominant in Romanies (13.7% and 10.3%, respectively). Y-DNA composition of Muslim Romani people from Šuto Orizari Municipality in North Macedonia, based on 57 samples: Haplogroup H – 59.6% Haplogroup E – 29.8% Haplogroup I – 5.3% Haplogroup R – 3.%, of which the half are R1b and many are R1a Haplogroup G – 1.8% Y-DNA Haplogroup H1a occurs in Romani at frequencies 7–70%. Unlike ethnic Hungarians, among Hungarian and Slovakian Romani subpopulations, Haplogroup E-M78 and I1 usually occur above 10% and sometimes over 20%. While among Slovakian and Tiszavasvari Romani the dominant haplogroup is H1a, among Tokaj Romani is Haplogroup J2a (23%), while among Taktaharkány Romani is Haplogroup I2a (21%). Five, rather consistent founder lineages throughout the subpopulations, were found among Romani – J-M67 and J-M92 (J2), H-M52 (H1a1), and I-P259 (I1). Haplogroup I-P259 as H is not found at frequencies of over 3 percent among host populations, while haplogroups E and I are absent in South Asia. The lineages E-V13, I-P37 (I2a) and R-M17 (R1a) may represent gene flow from the host populations. Bulgarian, Romanian and Greek Romani are dominated by Haplogroup H-M82 (H1a1), while among Spanish Romani J2 is prevalent. In Serbia among Kosovo and Belgrade Romani Haplogroup H prevails, while among Vojvodina Romani, H drops to 7 percent and E-V13 rises to a prevailing level. Among non-Roma Europeans Haplogroup H is extremely rare, peaking at 7 percent among Albanians from Tirana and 11 percent among Bulgarian Turks. It occurs at 5 percent among Hungarians, although the carriers might be of Romani origin. Among non Roma-speaking Europeans at 2