Pteranodon (); from Ancient Greek (pteron, "wing") and (anodon, "toothless") is a genus of pterosaur that included some of the largest known flying reptiles, with P. longiceps having a wingspan of over . They lived during the late Cretaceous geological period of North America in present-day Kansas, Nebraska, Wyoming, South Dakota and Alabama. More fossil specimens of Pteranodon have been found than any other pterosaur, with about 1,200 specimens known to science, many of them well preserved with nearly complete skulls and articulated skeletons. It was an important part of the animal community in the Western Interior Seaway. Pteranodon was not a dinosaur. By definition, all dinosaurs belong to the group Dinosauria; Pteranodon belongs to the group Pterosauria. Nonetheless, Pteranodon is the most famous pterosaur, frequently featured in dinosaur media and strongly associated with dinosaurs by the general public. While not dinosaurs, pterosaurs such as Pteranodon form a clade closely related to dinosaurs as both fall within the clade Avemetatarsalia. Discovery and history First fossils Pteranodon was the first pterosaur found outside of Europe. Its fossils first were found by Othniel Charles Marsh in 1871, in the Late Cretaceous Smoky Hill Chalk deposits of western Kansas. These chalk beds were deposited at the bottom of what was once the Western Interior Seaway, a large shallow sea over what now is the midsection of the North American continent. These first specimens, YPM 1160 and YPM 1161, consisted of partial wing bones, as well as a tooth from the prehistoric fish Xiphactinus, which Marsh mistakenly believed to belong to this new pterosaur (all known pterosaurs up to that point had teeth). In 1871, Marsh named the find Pterodactylus oweni, assigning it to the well-known (but much smaller) European genus Pterodactylus. Marsh also collected more wing bones of the large pterosaur in 1871. Realizing that the name he had chosen had already been used for Harry Seeley's European pterosaur species Pterodactylus oweni in 1864, Marsh renamed his giant North American pterosaur Pterodactylus occidentalis, meaning "Western wing finger," in his 1872 description of the new specimen. He named two additional species, based on size differences: Pterodactylus ingens (the largest specimen so far), and Pterodactylus velox (the smallest). Meanwhile, Marsh's rival Edward Drinker Cope had unearthed several specimens of the large North American pterosaur. Based on these specimens, Cope named two new species, Ornithochirus umbrosus and Ornithochirus harpyia, in an attempt to assign them to