(where stamens are present but their anthers do not produce viable pollen). Pollination of one flower by the pollen of a different flower of the same plant (geitonogamy) is common. The gynoecium consists of two carpels fused into a single, bicarpellate pistil with an inferior ovary. Stylopodia support two styles and secrete nectar, attracting pollinators like flies, mosquitoes, gnats, beetles, moths, and bees. The fruit is a schizocarp consisting of two fused carpels that separate at maturity into two mericarps, each containing a single seed. The fruits of many species are dispersed by wind but others such as those of Daucus spp., are covered in bristles, which may be hooked in sanicle Sanicula europaea and thus catch in the fur of animals. The seeds have an oily endosperm and often contain essential oils, containing aromatic compounds that are responsible for the flavour of commercially important umbelliferous seed such as anise, cumin and coriander. The shape and details of the ornamentation of the ripe fruits are important for identification to species level. Taxonomy Apiaceae was first described by John Lindley in 1836. The name is derived from the type genus Apium, which was originally used by Pliny the Elder circa 50 AD for a celery-like plant. The alternative name for the family, Umbelliferae, derives from the inflorescence being generally in the form of a compound umbel. The family was one of the first to be recognized as a distinct group in Jacques Daleschamps' 1586 Historia generalis plantarum. With Robert Morison's 1672 Plantarum umbelliferarum distribution nova it became the first group of plants for which a systematic study was published. The family is solidly placed within the Apiales order in the APG III system. It is closely related to Araliaceae and the boundaries between these families remain unclear. Traditionally groups within the family have been delimited largely based on fruit morphology, and the results from this have not been congruent with the more recent molecular phylogenetic analyses. The subfamilial and tribal classification for the family is currently in a state of flux, with many of the groups being found to be grossly paraphyletic or polyphyletic. Classification and phylogeny Prior to molecular phylogenetic studies, the family was subdivided primarily based on fruit characteristics. Molecular phylogenetic analyses from the mid-1990s onwards have shown that fruit characters evolved in parallel many times, so that using them in classification resulted in units that were not monophyletic.