noting that the feathers in the Munich specimen are poorly preserved. Nudds and Dyke reported a diameter of for the longest primary feather, which Paul could not confirm using photographs. Paul measured some of the inner primary feathers, finding rachises across. Despite these criticisms, Nudds and Dyke stood by their original conclusions. They claimed that Paul's statement, that an adult Archaeopteryx would have been a better flyer than the juvenile Munich specimen, was dubious. This, they reasoned, would require an even thicker rachis, evidence for which has not yet been presented. Another possibility is that they had not achieved true flight, but instead used their wings as aids for extra lift while running over water after the fashion of the basilisk lizard, which could explain their presence in lake and marine deposits (see Origin of avian flight). In 2004, scientists analysing a detailed CT scan of the braincase of the London Archaeopteryx concluded that its brain was significantly larger than that of most dinosaurs, indicating that it possessed the brain size necessary for flying. The overall brain anatomy was reconstructed using the scan. The reconstruction showed that the regions associated with vision took up nearly one-third of the brain. Other well-developed areas involved hearing and muscle coordination. The skull scan also revealed the structure of its inner ear. The structure more closely resembles that of modern birds than the inner ear of non-avian reptiles. These characteristics taken together suggest that Archaeopteryx had the keen sense of hearing, balance, spatial perception, and coordination needed to fly. Archaeopteryx had a cerebrum-to-brain-volume ratio 78% of the way to modern birds from the condition of non-coelurosaurian dinosaurs such as Carcharodontosaurus or Allosaurus, which had a crocodile-like anatomy of the brain and inner ear. Newer research shows that while the Archaeopteryx brain was more complex than that of more primitive theropods, it had a more generalized brain volume among Maniraptora dinosaurs, even smaller than that of other non-avian dinosaurs in several instances, which indicates the neurological development required for flight was already a common trait in the maniraptoran clade. Recent studies of flight feather barb geometry reveal that modern birds possess a larger barb angle in the trailing vane of the feather, whereas Archaeopteryx lacks this large barb angle, indicating potentially weak flight abilities.Archaeopteryx continues to play an important part in scientific debates about the origin and evolution of birds. Some scientists see it as