and A. siemensii, but their taxonomic history is complicated. Ten names have been published for the handful of specimens. As interpreted today, the name A. lithographica only referred to the single feather described by Meyer. In 1954 Gavin de Beer concluded that the London specimen was the holotype. In 1960, Swinton accordingly proposed that the name Archaeopteryx lithographica be placed on the official genera list making the alternative names Griphosaurus and Griphornis invalid. The ICZN, implicitly accepting De Beer's standpoint, did indeed suppress the plethora of alternative names initially proposed for the first skeleton specimens, which mainly resulted from the acrimonious dispute between Meyer and his opponent Johann Andreas Wagner (whose Griphosaurus problematicus – 'problematic riddle-lizard' – was a vitriolic sneer at Meyer's Archaeopteryx). In addition, in 1977, the Commission ruled that the first species name of the Haarlem specimen, crassipes, described by Meyer as a pterosaur before its true nature was realized, was not to be given preference over lithographica in instances where scientists considered them to represent the same species. It has been noted that the feather, the first specimen of Archaeopteryx described, does not correspond well with the flight-related feathers of Archaeopteryx. It certainly is a flight feather of a contemporary species, but its size and proportions indicate that it may belong to another, smaller species of feathered theropod, of which only this feather is known so far. As the feather had been designated the type specimen, the name Archaeopteryx should then no longer be applied to the skeletons, thus creating significant nomenclatorial confusion. In 2007, two sets of scientists therefore petitioned the ICZN requesting that the London specimen explicitly be made the type by designating it as the new holotype specimen, or neotype. This suggestion was upheld by the ICZN after four years of debate, and the London specimen was designated the neotype on 3 October 2011. Below is a cladogram published in 2013 by Godefroit et al. Species It has been argued that all the specimens belong to the same species, A. lithographica. Differences do exist among the specimens, and while some researchers regard these as due to the different ages of the specimens, some may be related to actual species diversity. In particular, the Munich, Eichstätt, Solnhofen, and Thermopolis specimens differ from the London, Berlin, and Haarlem specimens in being smaller or much larger, having different finger proportions, having more slender snouts lined with