either as a sister clade to Tubulidentata or as a grade leading to true tubulidentates. The first unambiguous tubulidentate was probably Myorycteropus africanus from Kenyan Miocene deposits. The earliest example from the genus Orycteropus was Orycteropus mauritanicus, found in Algeria in deposits from the middle Miocene, with an equally old version found in Kenya. Fossils from the aardvark have been dated to 5 million years, and have been located throughout Europe and the Near East. The mysterious Pleistocene Plesiorycteropus from Madagascar was originally thought to be a tubulidentate that was descended from ancestors that entered the island during the Eocene. However, a number of subtle anatomical differences coupled with recent molecular evidence now lead researchers to believe that Plesiorycteropus is a relative of golden moles and tenrecs that achieved an aardvark-like appearance and ecological niche through convergent evolution. Subspecies The aardvark has seventeen poorly defined subspecies listed: Orycteropus afer afer O. a. adametzi Grote, 1921 O. a. aethiopicus Sundevall, 1843 O. a. angolensis Zukowsky & Haltenorth, 1957 O. a. erikssoni Lönnberg, 1906 O. a. faradjius Hatt, 1932 O. a. haussanus Matschie, 1900 O. a. kordofanicus Rothschild, 1927 O. a. lademanni Grote, 1911 O. a. leptodon Hirst, 1906 O. a. matschiei Grote, 1921 O. a. observandus Grote, 1921 O. a. ruvanensis Grote, 1921 O. a. senegalensis Lesson, 1840 O. a. somalicus Lydekker, 1908 O. a. wardi Lydekker, 1908 O. a. wertheri Matschie, 1898 The 1911 Encyclopædia Britannica also mentions O. a. capensis or Cape ant-bear from South Africa. Description The aardvark is vaguely pig-like in appearance. Its body is stout with a prominently arched back and is sparsely covered with coarse hairs. The limbs are of moderate length, with the rear legs being longer than the forelegs. The front feet have lost the pollex (or 'thumb'), resulting in four toes, while the rear feet have all five toes. Each toe bears a large, robust nail which is somewhat flattened and shovel-like, and appears to be intermediate between a claw and a hoof. Whereas the aardvark is considered digitigrade, it appears at times to be plantigrade. This confusion happens because when it squats it stands on its soles. A contributing characteristic to the burrow digging capabilities of aardvarks is an endosteal tissue called compacted coarse cancellous bone (CCCB). The stress and strain resistance provided by CCCB allows aardvarks to create their burrows, ultimately leading to a favorable environment for plants and a