Ainu population. Recent autosomal evidence suggests that the Ainu derive a majority of their ancestry from the local Jōmon period people of Hokkaido. A 2019 study by Gakuhari et al., analyzing ancient Jōmon remains, finds about 79.3% Hokkaido Jōmon ancestry in the Ainu. Another 2019 study (by Kanazawa-Kiriyama et al.) finds about 66% Hokkaido Jōmon ancestry. A genetic study in 2021 (Sato et al.) found that the Ainu probably derived about ~49% of their ancestry from the local Hokkaido Jōmon, ~22% from the Okhotsk (samplified by Chukotko-Kamchatkan peoples), and ~29% from the Yamato Japanese. Population genomic data from various Jōmon period samples show that their main ancestry component split from other East Asian people at about 15,000 BCE. Following their migration into the Japanese archipelago, they became largely isolated from outside geneflow. However geneflow from Ancient North Eurasians towards the Jōmon period population was detected along a North to South cline, with a peak among Hokkaido Jōmon. Physical description Physical differences could be observed between different Ainu subgroups and clans. According to anthropologists "…features considered to distinguish the Ainu from other populations in the area, especially the Japanese, are the tendency to dolichocephaly (long-headedness), a well developed glabella, a deeply depressed nose root, widely projecting cheekbones, a comparatively massive mandible (lower jaw), and an edge to edge bite", as well as more body and facial hair. Many Ainu men have abundant wavy hair and often wear long beards. The book of Ainu Life and Legends by author Kyōsuke Kindaichi (published by the Japanese Tourist Board in 1942) contains a physical description of Ainu: Many have wavy hair, but some straight black hair. Very few of them have wavy brownish hair. Their skins are generally reported to be light brown. But this is due to the fact that they labor on the sea and in briny winds all day. Old people who have long desisted from their outdoor work are often found to be as white as western men. The Ainu have broad faces, beetling eyebrows, and sometimes large sunken eyes, which are generally horizontal and of the so-called European type. Eyes of the Mongolian type are rare but occasionally found among them. A comparative study by Brace et al. (2001) argues for a closer morphological relation of the Ainu with prehistoric and living European groups, compared to with other East Asian groups. The authors concluded that part of their ancestors