a sphere of gas held up against gravity by internal thermal pressure, and one of Eddington's chief additions was to show that radiation pressure was necessary to prevent collapse of the sphere. He developed his model despite knowingly lacking firm foundations for understanding opacity and energy generation in the stellar interior. However, his results allowed for calculation of temperature, density and pressure at all points inside a star (thermodynamic anisotropy), and Eddington argued that his theory was so useful for further astrophysical investigation that it should be retained despite not being based on completely accepted physics. James Jeans contributed the important suggestion that stellar matter would certainly be ionized, but that was the end of any collaboration between the pair, who became famous for their lively debates. Eddington defended his method by pointing to the utility of his results, particularly his important mass–luminosity relation. This had the unexpected result of showing that virtually all stars, including giants and dwarfs, behaved as ideal gases. In the process of developing his stellar models, he sought to overturn current thinking about the sources of stellar energy. Jeans and others defended the Kelvin–Helmholtz mechanism, which was based on classical mechanics, while Eddington speculated broadly about the qualitative and quantitative consequences of possible proton–electron annihilation and nuclear fusion processes. Around 1920, he anticipated the discovery and mechanism of nuclear fusion processes in stars, in his paper "The Internal Constitution of the Stars". At that time, the source of stellar energy was a complete mystery; Eddington correctly speculated that the source was fusion of hydrogen into helium, liberating enormous energy according to Einstein's equation . This was a particularly remarkable development since at that time fusion and thermonuclear energy, and even the fact that stars are largely composed of hydrogen (see metallicity), had not yet been discovered. Eddington's paper, based on knowledge at the time, reasoned that: The leading theory of stellar energy, the contraction hypothesis (cf. the Kelvin–Helmholtz mechanism), should cause stars' rotation to visibly speed up due to conservation of angular momentum. But observations of Cepheid variable stars showed this was not happening. The only other known plausible source of energy was conversion of matter to energy; Einstein had shown some years earlier that a small amount of matter was equivalent to a large amount of energy. Francis Aston had also recently shown that the mass of a helium atom was about 0.8% less