valuable energy while still allowing the Tyrannosaurus to actively chase down and kill its intended target. Gregory S. Paul stated that tyrannosaurids attacked hadrosaurs by targeting the caudofemoralis muscle and thigh muscle to disable their prey's locomotor system, the abdomen to eviscerate their prey and the neck which would have caused the most rapid death by damaging the trachea and major blood vessels. Tyrannosaurus was once proposed to have had infectious saliva used to kill its prey. This theory was first proposed by William Abler. Abler examined the teeth of tyrannosaurids between each tooth serration; the serrations may have held pieces of carcass with bacteria, giving Tyrannosaurus a deadly, infectious bite much like the Komodo dragon was originally thought to have. However, Jack Horner regards Tyrannosaurus tooth serrations as more like cubes in shape than the serrations on a Komodo monitor's teeth, which are rounded. All forms of saliva contain possibly hazardous bacteria, so the prospect of it being used as a method of predation is disputable. Feeding behavior A few studies have also focused on Tyrannosaurus complex feeding habits. Tyrannosaurus, and most other theropods, probably primarily processed carcasses with lateral shakes of the head, like crocodilians. The head was not as maneuverable as the skulls of allosauroids, due to flat joints of the neck vertebrae. Eric Snively and Anthony Russell further elaborates in a study published in 2007 that Tyrannosaurus had a powerful neck that would have enabled it to strike rapidly at prey and take on complex and modulated inertial feeding; a way of feeding used by modern archosaurs that involved the animal ripping away chunks of meat, tossing it into the air and swallowing it. A team of paleontologists, led by Denver Fowler from the University of the Rockies would later discover that Tyrannosaurus employed a complex feeding strategy to consume Triceratops after analyzing various specimens. This involved the theropod repositioning and tearing off the head of the dead Triceratops, so that it could consume its meal's nutrient-rich neck muscles. Studies of Tyrannosauruss teeth have also revealed that the animal's dentition was more complex than previously imagined. Researchers Kirstin Brink, Robert Reisz et al. found that Tyrannosaurus and other carnivorous theropods were equipped with teeth that were highly complex for carnivores; specialized layers of dentine enlarged the serration on the inside of the tooth structure. This unique structure, found in other large predators from other prehistoric eras,