with each new pair only becoming functional at the next moult. After the final furcilia stage, the krill resembles the adult. Decapoda Apart from the prawns of the suborder Dendrobranchiata, all decapod crustaceans brood their eggs on the female's pleopods. This has resulted in development in decapod crustaceans being generally abbreviated. There are at most nine larval stages in decapods, as in krill, and both decapod nauplii and krill nauplii often lack mouthparts and survive on nutrients supplied in the egg yolk (lecithotrophy). In species with normal development, eggs are roughly 1% of the size of the adult; in species with abbreviated development, and therefore more yolk in the eggs, the eggs may reach 1/9 of the adult's size. The post-larva of shrimp is called parva, after the species Acanthephyra parva described by Henri Coutière, but which was later recognised as the larva of Acanthephyra purpurea. In the marine lobsters, there are three larval stages, all similar in appearance. Freshwater crayfish embryos differ from those of other crustaceans in having 40 ectoteloblast cells, rather than around 19. The larvae show abbreviated development, and hatch with a full complement of adult appendages with the exceptions of the uropods and the first pair of pleopods. The larvae of the Achelata (slipper lobsters, spiny lobsters and furry lobsters) are unlike any other crustacean larvae. The larvae are known as phyllosoma, after the genus Phyllosoma erected by William Elford Leach in 1817. They are flattened and transparent, with long legs and eyes on long eyestalks. After passing through 8–10 phyllosoma stages, the larva undergoes "the most profound transformation at a single moult in the Decapoda", when it develops into the so-called puerulus stage, which is an immature form resembling the adult animal. The members of the traditional infraorder Thalassinidea can be divided into two groups on the basis of their larvae. According to Robert Gurney, the "homarine group" comprises the families Axiidae and Callianassidae, while the "anomuran group" comprises the families Laomediidae and Upogebiidae. This split corresponds with the division later confirmed with molecular phylogenetics. Among the Anomura, there is considerable variation in the number of larval stages. In the South American freshwater genus Aegla, the young hatch from the eggs in the adult form. Squat lobsters pass through four, or occasionally five, larval states, which have a long rostrum, and a spine on either side of the carapace; the first post-larva closely resembles