to revised estimates, and "Alexander (1996) argued that based on the bone dimensions of Tyrannosaurus it is unlikely they could have travelled at more than 8ms−1." Several calculations using variants of the formula indicate that dinosaurs probably travelled at around 3 ms−1 with a top speed of 8 ms−1. This translates to a speed range of roughly 6–20 mph. Alexander was secretary of the Zoological Society of London (1992–1999) which included supervising the management of London and Whipsnade Zoos. He was president of the Society for Experimental Biology (1995–1997), President of the International Society of Vertebrate Morphologists (1997–2001) and editor of the Proceedings of the Royal Society B (1998–2004). Alexander specialised in research on animal mechanics and published numerous books and research papers in the field from 1959. Film and TV work Horizon (1976) TV series documentary The Hot-Blooded Dinosaurs (1976) The Dinosaurs! (1992) Walking with Beasts (2001) TV series documentary (principal scientific advisor) The Future Is Wild (2003) TV series documentary Extinct: A Horizon Guide to Dinosaurs (2001) TV documentary Honours and awards Alexander received several awards and honours during his career including: 1979 Linnean Medal for Zoology Linnean Society of London 1987 Elected a Fellow of the Royal Society (FRS) 2000 Birthday Honours List Commander of the Most Excellent Order of the British Empire (CBE) 2001 Foreign Honorary Member American Academy of Arts and Sciences 2002 Honorary Fellow Zoological Society of London 2003 Borelli Award American Society of Biomechanics Personal life Alexander married Ann Elizabeth Coulton in 1961. They had a son and a daughter. Death Alexander died in 2016, aged 81. He was survived by his wife and children. Selected publications Books Functional Design in Fishes, Hutchinson University Library, 1967, 1970 Animal Mechanics, Sidgwick & Jackson, 1968 Size and Shape, Edward Arnold, 1971 The Chordates, Cambridge University Press, 1975 Mechanics and energetics of animal locomotion, with G. Goldspink, Halsted Press, 1977 The Invertebrates, Cambridge University Press, 1979 Optima for Animals, Hodder Arnold, 1982 Locomotion of animals, Springer, 1985 The Collins Encyclopedia of Animal Biology, HarperCollins Publishers, 1986 Elastic Mechanisms in Animal Movement, Cambridge University Press, 1988 Dynamics of Dinosaurs and other Extinct Giants, Columbia University Press, 1989 Animals, Cambridge University Press, 1990 How dinosaurs ran, Scientific American, 1991 Animals, Cambridge University Press, 1991 The Human Machine, Natural History Museum, Stationery Office Books, 1992 Exploring Biomechanics: Animals in Motion, W H Freeman & Co, 1992 Bones: The Unity