possibly Megaladapis. Flacourt described it as: Local tales of a (Malagasy for 'cow that is not a cow'), or pygmy hippopotamus, led French naturalist Alfred Grandidier to follow a village headman to a marsh in southwestern Madagascar, a site called Ambolisatra, which became the first known subfossil site in Madagascar. In 1868, Grandidier uncovered the first subfossil remains of lemurs—a humerus from Palaeopropithecus and a tibia of a sifaka. The Palaeopropithecus remains were not described for several decades, and it took decades more for the remains to be correctly paired with other sloth lemur remains. It was not until 1893 that giant lemur species were formally described, when Charles Immanuel Forsyth Major discovered and described a long, narrow skull of Megaladapis madagascariensis in a marsh. His discoveries in various marshes of central and southwestern Madagascar sparked paleontological interest, resulting in an overabundance of taxonomic names and confused assemblages of bones from numerous species, including non-primates. Specimens were distributed between European museums and Madagascar, often resulting in the loss of field data that went with the specimens, if the data had been recorded at all. In 1905, Alfred Grandidier's son, Guillaume Grandidier, reviewed subfossil lemur taxonomy and determined that too many names had been created. His review established most of the presently known family and genera names for the extinct lemurs. Despite the taxonomic clarification, subfossil postcrania from different genera, particularly Megaladapis, Palaeopropithecus and Hadropithecus, continued to be incorrectly paired and sometimes assigned to non-primates. Since subfossil remains were often dredged from marshes one by one, pairing skulls with other bones was often guesswork based on size-matching, and was not very accurate as a consequence. Even as late as the 1950s, bones of non-primates were attributed to subfossil lemurs. One reconstruction of the confounded subfossil remains by paleontologist Herbert F. Standing depicted Palaeopropithecus as an aquatic animal that swam near the surface, keeping its eyes, ears, and nostrils slightly above water. Postcranial remains of Palaeopropithecus had previously been paired with Megaladapis by Guillaume Grandidier, who viewed it as a giant tree sloth, which he named Bradytherium. Standing's aquatic theory was supported by Italian paleontologist Giuseppe Sera, who reconstructed Palaeopropithecus as an "arboreal-aquatic acrobat" that not only swam in water but climbed trees and dove from there into the water. Sera took the aquatic theory further in 1938 by including other extinct lemurs, including Megaladapis, which he viewed as a thin ray-like