and Nod and M.K. share a kiss before M.K. is returned to her original size. M.K. reunites with Bomba and later becomes his assistant; the two of them remain in contact with their small friends as they continue the research of their world. Voice cast Colin Farrell as Ronin, a seasoned Leafman warrior, leader of the Leafmen, Nod's guardian and teacher, and a friend of Nod's late father. Josh Hutcherson as Nod, a rookie Leafman warrior, Ronin's student, and M.K.'s love interest and boyfriend. Amanda Seyfried as Mary Katherine "M.K." Bomba, Professor Bomba's 17-year-old daughter. She is named after William Joyce's late daughter. Christoph Waltz as Mandrake, the cruel and snide owner of the Boggans. Waltz also voiced Mandrake in the German dub of the film since German is his native language. Aziz Ansari as Mub, a slug, Grub's best friend and caretaker of the pods. Chris O'Dowd as Grub, a snail, Mub's best friend and fellow caretaker of the pods. Pitbull as Bufo, a toad who works as a race fixer and a businessman. Jason Sudeikis as Professor Radcliffe Bomba, M.K.'s father and a scientist. Steven Tyler as Nim Galuu, a Glowworm, a showman and keeper of magic scrolls that tell what has occurred during the times, in a tree. Beyoncé Knowles as Queen Tara, the Mother Nature-like queen of the forest and Ronin's childhood love. Blake Anderson as Dagda, Mandrake's son and the Boggan's general officer. After his death, Mandrake is intent on destroying the forest. Judah Friedlander as Larry, a taxi driver. Thomas F. Wilson as Finn, second-in-command of the Leafmen. His name only appears in the closing credits. John DiMaggio as Pinecone Jinn. Rosa Salazar as Roller Derby Girl. Kyle Kinane as Biker Dude. Emma Kenney as Marigold Girl. Production In 2006, it was reported that Chris Wedge would be directing an animated feature film based on William Joyce's book, The Leaf Men and the Brave Good Bugs for Fox Animation. Joyce, who had already collaborated with Wedge as a designer and producer on the 2005 film Robots, was set to produce the film. At one point, Wedge got permission to find a new home for the film and turned to Pixar, led by John Lasseter, a close friend that Wedge knew from working on Tron. When Pixar tried to close the rights for the film and start development, Fox changed their mind, and the film